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Source: Checklist of the Scarabaeoidea of the Nearctic Realm, website (version 4)

ITIS records citing this source


Kingdom Animalia
	Abcrana Saylor, 1942 -- invalid
	Acanthocerus MacLeay, 1819 -- invalid
	Acanthocerus laevistriatus Laporte, 1840 -- invalid
	Acanthurus Kirby, 1827 -- invalid
	Aclinidia Casey, 1915 -- invalid
	Acoma Casey, 1889 -- valid
	Acoma arizonica Brown, 1929 -- valid
	Acoma brunnea Casey, 1889 -- valid
	Acoma cazieri Saylor, 1948 -- valid
	Acoma confusa Van Dyke, 1928 -- valid
	Acoma conjuncta Howden, 1962 -- valid
	Acoma dilemma Saylor, 1948 -- valid
	Acoma diminiata Howden, 1958 -- valid
	Acoma evansi Howden, 1962 -- valid
	Acoma gibsoni Howden, 1962 -- valid
	Acoma glabrata Cazier, 1953 -- valid
	Acoma granulifrons Howden, 1958 -- valid
	Acoma incognita Howden, 1958 -- valid
	Acoma knulli Howden, 1958 -- valid
	Acoma leechi Cazier, 1953 -- valid
	Acoma martini Howden, 1962 -- valid
	Acoma mimica Howden, 1962 -- valid
	Acoma minuta Cazier, 1953 -- valid
	Acoma mixta Howden, 1958 -- valid
	Acoma nigrita Cazier, 1953 -- invalid
	Acoma ochlera Howden, 1958 -- valid
	Acoma parva Howden, 1958 -- valid
	Acoma robusta Van Dyke, 1928 -- valid
	Acoma rossi Saylor, 1948 -- valid
	Acoma rufula Howden, 1958 -- valid
	Acoma seticollis Howden, 1958 -- valid
	Acoma sexfoliata Saylor, 1948 -- valid
	Acoma stathamae Cazier, 1953 -- valid
	Acratus Horn, 1867 -- invalid
	Acrossus Mulsant, 1842 -- valid
	Acrossus rubripennis (Horn, 1870) -- valid
	Acrossus rufipes (Linnaeus, 1758) -- valid
	Aegialia Latreille, 1807 -- valid
	Aegialia (Aegialia) Latreille, 1807 -- valid
	Aegialia (Psammoporus) Thomson, 1863 -- valid
	Aegialia amplipunctata Gordon and Cartwright, 1988 -- valid
	Aegialia arenaria (Fabricius, 1787) -- valid
	Aegialia blanchardi Horn, 1887 -- valid
	Aegialia carri Gordon and Cartwright, 1988 -- valid
	Aegialia cartwrighti Stebnicka, 1977 -- valid
	Aegialia concinna Gordon and Cartwright, 1977 -- valid -- Ciervo aegialian scarab
	Aegialia conferta Horn, 1871 -- valid
	Aegialia convexa Fall, 1932 -- valid
	Aegialia crassa LeConte, 1857 -- valid
	Aegialia crescenta Gordon and Cartwright, 1977 -- valid -- Crescent Dunes aegialian scarab
	Aegialia criddlei Brown, 1931 -- valid
	Aegialia cylindrica (Eschscholtz, 1822) -- valid
	Aegialia exarata Mannerheim, 1853 -- valid
	Aegialia hardyi Gordon and Cartwright, 1977 -- valid -- Hardy's aegialian scarab
	Aegialia insularis Brown, 1931 -- invalid
	Aegialia kelsoi Gordon and Cartwright, 1988 -- valid
	Aegialia knighti Gordon and Rust, 1997 -- valid
	Aegialia lacustris LeConte, 1850 -- valid
	Aegialia latispina LeConte, 1878 -- valid
	Aegialia magnifica Gordon and Cartwright, 1977 -- valid -- large aegialian scarab
	Aegialia mcclevei Gordon, 1990 -- valid
	Aegialia nana Brown, 1931 -- valid
	Aegialia nigrella Brown, 1931 -- valid
	Aegialia opaca Brown, 1931 -- valid
	Aegialia opifex Horn, 1887 -- valid
	Aegialia punctata Brown, 1931 -- valid
	Aegialia rufa LeConte, 1878 -- invalid
	Aegialia rufina Silfverberg, 1977 -- invalid
	Aegialia spinosa Gordon and Cartwright, 1988 -- valid
	Aegialia spissipes LeConte, 1878 -- invalid
	Aegialia terminalis Brown, 1931 -- valid
	Aegialiini Laporte, 1840 -- valid
	Aesalinae MacLeay, 1819 -- valid
	Afroglaresis Petrovitz, 1968 -- invalid
	Aglycoptera Sharp, 1885 -- invalid
	Agoliinus Schmidt, 1913 -- valid
	Agoliinus albertanus (Brown, 1928) -- valid
	Agoliinus aleutus (Eschscholtz, 1822) -- valid
	Agoliinus anthracus Gordon and Skelley, 2007 -- valid
	Agoliinus aquilonarius (Brown, 1928) -- valid
	Agoliinus ashworthi (Gordon, 2006) -- valid
	Agoliinus bidentatus (Schmidt, 1907) -- valid
	Agoliinus canadensis (Garnett, 1920) -- valid
	Agoliinus congregatus (Mannerheim, 1853) -- valid
	Agoliinus corruptor (Brown, 1929) -- valid
	Agoliinus cruentatus (LeConte, 1878) -- valid
	Agoliinus explanatus (LeConte, 1878) -- valid
	Agoliinus guttatus (Eschscholtz, 1823) -- valid
	Agoliinus hatchi Gordon and Skelley, 2007 -- valid
	Agoliinus idahoensis Gordon and Skelley, 2007 -- valid
	Agoliinus incommunis (Fall, 1932) -- valid
	Agoliinus ingenursus Gordon and Skelley, 2007 -- valid
	Agoliinus leopardus (Horn, 1870) -- valid
	Agoliinus malkini (Hatch, 1971) -- valid
	Agoliinus manitobensis (Brown, 1928) -- valid
	Agoliinus piceatus (Robinson, 1946) -- valid
	Agoliinus plutonicus (Fall, 1907) -- valid
	Agoliinus poudreus Gordon and Skelley, 2007 -- valid
	Agoliinus praealtus Gordon and Skelley, 2007 -- valid
	Agoliinus sigmoideus (Van Dyke, 1918) -- valid
	Agoliinus wickhami (Brown, 1928) -- valid
	Agrilinellus Dellacasa, Dellacasa and Gordon, 2008 -- valid
	Agrilinellus nuriae Dellacasa, Dellacasa and Gordon, 2008 -- valid
	Agrilinellus ornatus (Schmidt, 1911) -- valid
	Aidophus Balthasar, 1963 -- valid
	Aidophus parcus (Horn, 1887) -- valid
	Aidophus skelleyi Harpootlian and Gordon, 2002 -- valid
	Alamona Casey, 1915 -- invalid
	Alamona parviceps Casey, 1915 -- invalid
	Albridarollia Bol�var y Pieltain, Jim�nez-As�a and Mart�nez, 1963 -- invalid
	Alobus LeConte, 1856 -- invalid
	Alocerus Zang, 1905 -- invalid
	Amblonoxia Reitter, 1902 -- valid
	Amblonoxia bidentata (Fall, 1932) -- valid
	Amblonoxia carpenteri (LeConte, 1876) -- valid
	Amblonoxia fieldi (Fall, 1908) -- valid
	Amblonoxia harfordi (Casey, 1889) -- valid
	Amblonoxia palpalis (Horn, 1880) -- valid
	Amblonoxia riversi (Casey, 1895) -- valid
	Amblyphileurus Kolbe, 1910 -- invalid
	Amechanus Horn, 1870 -- invalid
	Amidorinus Koshantschikov, 1912 -- invalid
	Amphimallon Latreille, 1825 -- valid
	Amphimallon korbi Reitter, 1894 -- invalid
	Amphimallon majale (Razumowski, 1789) -- valid
	Amphimallon pellitulum Reitter, 1902 -- invalid
	Amphimallum Agassiz, 1847 -- invalid
	Anaeretes Dejean, 1836 -- invalid
	Anagryllus Casey, 1915 -- invalid
	Anamnesis Vigors, 1826 -- invalid
	Anastrategus Casey, 1915 -- invalid
	Anastrategus carolinensis Casey, 1915 -- invalid
	Anastrategus cavicauda Casey, 1915 -- invalid
	Anastrategus cognatus Casey, 1915 -- invalid
	Anastrategus durangoensis Casey, 1915 -- invalid
	Anastrategus inflatus Casey, 1915 -- invalid
	Anastrategus tantalus Casey, 1915 -- invalid
	Anatropis Casey, 1915 -- invalid
	Ancognatha Erichson, 1847 -- valid
	Ancognatha aequata Bates, 1888 -- invalid
	Ancognatha durangoana Casey, 1915 -- invalid
	Ancognatha laevigata Bates, 1888 -- invalid
	Ancognatha manca (LeConte, 1866) -- valid
	Ancognatha perspicua Casey, 1915 -- invalid
	Ancognatha zuniella Casey, 1915 -- invalid
	Ancylonycha brevicollis Blanchard, 1851 -- invalid
	Ancylonycha cognata Burmeister, 1855 -- invalid
	Ancylonycha comata Burmeister, 1855 -- invalid
	Ancylonycha consequens Walker, 1866 -- invalid
	Ancylonycha cribricollis Blanchard, 1851 -- invalid
	Ancylonycha fimbriata Burmeister, 1855 -- invalid
	Ancylonycha gibbosa Burmeister, 1855 -- invalid
	Ancylonycha guadulpensis Blanchard, 1851 -- invalid
	Ancylonycha laticeps Blanchard, 1851 -- invalid
	Ancylonycha longicornis Burmeister, 1855 -- invalid
	Ancylonycha nigropicea Walker, 1866 -- invalid
	Ancylonycha pruinosa Melsheimer, 1845 -- invalid
	Ancylonycha puncticollis Blanchard, 1851 -- invalid
	Ancylonycha uninotata Walker, 1866 -- invalid
	Anisophileurus Prell, 1914 -- invalid
	Anisopliina Burmeister, 1844 -- valid
	Annegialia Howden, 1971 -- valid
	Annegialia ataeniformis Howden, 1971 -- valid
	Anomala Samouelle, 1819 -- valid
	Anomala adscita (Robinson, 1941) -- valid
	Anomala alterata Arrow, 1915 -- invalid
	Anomala amissa Casey, 1915 -- invalid
	Anomala antennata Schaeffer, 1906 -- valid
	Anomala apacheana Wickham, 1913 -- invalid
	Anomala arida Casey, 1915 -- valid
	Anomala beckeri Ohaus, 1897 -- valid
	Anomala binotata (Gyllenhal, 1817) -- valid
	Anomala butleri (Howden, 1955) -- valid
	Anomala camancha Wickham, 1913 -- invalid
	Anomala canadensis Casey, 1915 -- invalid
	Anomala carinifrons Bates, 1888 -- valid
	Anomala carlsoni Hardy, 1976 -- valid
	Anomala castaniceps Bates, 1888 -- valid
	Anomala cavifrons LeConte, 1867 -- valid
	Anomala centralis LeConte, 1863 -- valid
	Anomala chevrolati Bates, 1888 -- valid
	Anomala cincta Say, 1835 -- valid
	Anomala cincta cincta Say, 1835 -- valid
	Anomala cinctella Burmeister, 1844 -- invalid
	Anomala compacta Casey, 1915 -- invalid
	Anomala crinicollis Ohaus, 1902 -- valid
	Anomala delicata Casey, 1915 -- valid
	Anomala diabla Potts, 1976 -- valid
	Anomala dichroa Melsheimer, 1845 -- invalid
	Anomala digressa Casey, 1915 -- valid
	Anomala discoidalis Bates, 1888 -- valid
	Anomala dubia (Scopoli, 1763) -- valid
	Anomala dubia Blatchley, 1910 -- invalid
	Anomala ellipsis Casey, 1915 -- valid
	Anomala exigua (Schwarz, 1878) -- valid -- Exiguous anomalan scarab
	Anomala eximia Potts, 1976 -- valid -- Archbold anomalan scarab
	Anomala flavilla Bates, 1888 -- valid
	Anomala flavilla coachellae Potts, 1977 -- valid
	Anomala flavilla flavilla Bates, 1888 -- valid
	Anomala flavipennis Burmeister, 1844 -- valid
	Anomala flavipennis aransas Potts, 1977 -- valid
	Anomala flavipennis flavipennis Burmeister, 1844 -- valid
	Anomala flavipennis okaloosensis Potts, 1977 -- valid
	Anomala flavipennis subquadrata Casey, 1915 -- valid
	Anomala flohri Ohaus, 1897 -- valid
	Anomala floridana Casey, 1915 -- invalid
	Anomala foraminosa Bates, 1888 -- valid
	Anomala hardyorum Potts, 1976 -- valid -- Hardy's dune beetle
	Anomala histrionella Bates, 1888 -- valid
	Anomala imperialae Potts, 1976 -- valid
	Anomala innuba (Fabricius, 1787) -- valid
	Anomala insitiva Robinson, 1938 -- valid
	Anomala kanei Potts, 1976 -- valid
	Anomala kansana Hayes and McColloch, 1924 -- invalid
	Anomala latreillei Blanchard, 1851 -- invalid
	Anomala lucicola (Fabricius, 1798) -- valid
	Anomala ludoviciana Schaeffer, 1906 -- valid
	Anomala luteipennis LeConte, 1854 -- invalid
	Anomala marginata (Fabricius, 1793) -- valid
	Anomala marginella LeConte, 1854 -- invalid
	Anomala medorensis Casey, 1915 -- invalid
	Anomala mendica Casey, 1915 -- valid
	Anomala minuta Burmeister, 1844 -- invalid
	Anomala modulata Casey, 1915 -- invalid
	Anomala moquina Casey, 1915 -- invalid
	Anomala nigropicta Casey, 1915 -- invalid
	Anomala nimbosa Casey, 1915 -- valid
	Anomala oblivia Horn, 1884 -- valid
	Anomala orientalis (Waterhouse, 1875) -- valid
	Anomala papagoana Casey, 1915 -- invalid
	Anomala parvula Burmeister, 1844 -- valid
	Anomala peninsularis Schaeffer, 1906 -- valid
	Anomala phylloperthoides Nonfried, 1894 -- invalid
	Anomala piceola Casey, 1915 -- invalid
	Anomala pinicola Melsheimer, 1845 -- invalid
	Anomala pubescens Blatchley, 1910 -- invalid
	Anomala raydoma Saylor, 1948 -- invalid
	Anomala robinsoni Potts, 1974 -- valid
	Anomala sabinae Potts, 1976 -- valid
	Anomala sagax Casey, 1915 -- invalid
	Anomala saginulata Casey, 1915 -- invalid
	Anomala semilivida LeConte, 1878 -- valid
	Anomala servilis Casey, 1915 -- invalid
	Anomala stigmatella Casey, 1915 -- invalid
	Anomala suavis Potts, 1976 -- valid
	Anomala tenera Casey, 1915 -- invalid
	Anomala tibialis Schaeffer, 1906 -- valid -- Tibial scarab
	Anomala umbra Casey, 1915 -- valid
	Anomala undulata Melsheimer, 1845 -- valid
	Anomala varians Burmeister, 1844 -- invalid
	Anomalacra Casey, 1915 -- valid
	Anomalacra clypealis (Schaeffer, 1907) -- valid
	Anomalacra cuneata Casey, 1915 -- invalid
	Anomalacra werneri Howden, 1956 -- invalid
	Anomalaegialia Brown, 1931 -- invalid
	Anomalepta Casey, 1915 -- invalid
	Anomalepta flaccida Casey, 1915 -- invalid
	Anomalina Streubel, 1839 -- valid
	Anomalini Streubel, 1839 -- valid
	Anomalopides Strand, 1928 -- invalid
	Anomalopus Casey, 1915 -- invalid
	Anoplocephalus Schaeffer, 1906 -- invalid
	Anoplognatho Rivers, 1888 -- valid
	Anoplognatho dunnianus Rivers, 1888 -- valid
	Apeltastes Howden, 1968 -- valid
	Apeltastes elongatus Howden, 1968 -- valid
	Aphodiinae Leach, 1815 -- valid
	Aphodiini Leach, 1815 -- valid
	Aphodinus Motschulsky, 1862 -- invalid
	Aphodius Illiger, 1798 -- valid
	Aphodius adelaidae Hope, 1847 -- invalid
	Aphodius agraphus Mader, 1953 -- invalid
	Aphodius ambiguus Boheman, 1858 -- invalid
	Aphodius amplinotum Gordon and Howden, 1973 -- invalid
	Aphodius angularis LeConte, 1850 -- invalid
	Aphodius angustatus Mulsant, 1842 -- invalid
	Aphodius angustus Hatch, 1971 -- invalid
	Aphodius angustus Philippi and Philippi, 1864 -- invalid
	Aphodius anthracinus LeConte, 1878 -- invalid
	Aphodius anxius Mulsant, 1842 -- invalid
	Aphodius arcticus Harold, 1863 -- invalid
	Aphodius aterrimus Melsheimer, 1845 -- invalid
	Aphodius atwateri Cartwright, 1944 -- invalid
	Aphodius auctus Mulsant, 1842 -- invalid
	Aphodius aurelianus Harold, 1863 -- invalid
	Aphodius awemeanus Brown, 1928 -- invalid
	Aphodius baseolus Mulsant, 1842 -- invalid
	Aphodius bidens LeConte, 1878 -- invalid
	Aphodius biformis Reitter, 1892 -- invalid
	Aphodius bifrons Bates, 1887 -- invalid
	Aphodius binominatus Schmidt, 1907 -- invalid
	Aphodius bistrigatus Dalla Torre, 1879 -- invalid
	Aphodius blaisdelli Fall, 1905 -- invalid
	Aphodius boum Gistel, 1857 -- invalid
	Aphodius brevitarsis P�ringuey, 1901 -- invalid
	Aphodius brunnescens Reitter, 1892 -- invalid
	Aphodius calgaryensis Balthasar, 1970 -- invalid
	Aphodius campestris Blatchley, 1912 -- invalid
	Aphodius capicola Harold, 1862 -- invalid
	Aphodius capraram Gistel, 1857 -- invalid
	Aphodius carbonarius Sturm, 1805 -- invalid
	Aphodius caseyi Saylor, 1940 -- invalid
	Aphodius castaneipennis Waterhouse, 1875 -- invalid
	Aphodius catulus Balthasar, 1946 -- invalid
	Aphodius centrolineatoides Strand and Gulbis, 1957 -- invalid
	Aphodius cincticulus Hope, 1847 -- invalid
	Aphodius cinereus Mulsant and Rey, 1870 -- invalid
	Aphodius cockerelli Leng, 1918 -- invalid
	Aphodius columbiensis Wickham, 1913 -- invalid
	Aphodius comanchi Robinson, 1940 -- invalid
	Aphodius concolor Mulsant, 1842 -- invalid
	Aphodius confluens Schilsky, 1888 -- invalid
	Aphodius constrictus Robinson, 1946 -- invalid
	Aphodius controversus Balthasar, 1932 -- invalid
	Aphodius copronymus Melsheimer, 1845 -- invalid
	Aphodius corvinus Haldeman, 1848 -- invalid
	Aphodius cressoni Robinson, 1946 -- invalid
	Aphodius cribratus Mulsant, 1842 -- invalid
	Aphodius cribratus LeConte, 1878 -- invalid
	Aphodius cuniculus Chevrolat, 1864 -- invalid
	Aphodius curtus Haldeman, 1848 -- invalid
	Aphodius davisi Fall, 1930 -- invalid
	Aphodius diabolicus Balthasar, 1932 -- invalid
	Aphodius discoidalis Gistel, 1857 -- invalid
	Aphodius discus Wiedemann, 1823 -- invalid
	Aphodius distinctoides Strand and Gulbis, 1957 -- invalid
	Aphodius diversus Waterhouse, 1875 -- invalid
	Aphodius edai Balthasar, 1956 -- invalid
	Aphodius elongatus M�n�tries, 1832 -- invalid
	Aphodius emarginatus Stephens, 1830 -- invalid
	Aphodius errans Brown, 1930 -- invalid
	Aphodius expertus Harold, 1871 -- invalid
	Aphodius explanatus Brown, 1928 -- invalid
	Aphodius fenyesi Van Dyke, 1947 -- invalid
	Aphodius fimetarius (Linnaeus, 1758) -- valid
	Aphodius flavocinctus Harold, 1860 -- invalid
	Aphodius flavogriseus Mulsant, 1842 -- invalid
	Aphodius flohri Bates, 1887 -- invalid
	Aphodius fordi Gordon, 1974 -- invalid -- Ford's aphodius scarab
	Aphodius fortunus Fall, 1932 -- invalid
	Aphodius freyi Balthasar, 1941 -- invalid
	Aphodius fumosus Mulsant, 1842 -- invalid
	Aphodius garmani Brown, 1929 -- invalid
	Aphodius gentneri Hatch, 1971 -- invalid
	Aphodius gisteli Strand, 1917 -- invalid
	Aphodius goffi Cartwright, 1939 -- invalid
	Aphodius gonagricus Fischer von Waldheim, 1844 -- invalid
	Aphodius griseolus Mulsant, 1842 -- invalid
	Aphodius griseus Dalla Torre, 1879 -- invalid
	Aphodius guadeloupensis Fleutiaux and Sall�, 1889 -- invalid
	Aphodius haemorrhous Stephens, 1830 -- invalid
	Aphodius haywardi Brown, 1928 -- invalid
	Aphodius hemicyclus Mulsant, 1842 -- invalid
	Aphodius humeralis B�guena, 1927 -- invalid
	Aphodius humeralis LeConte, 1878 -- invalid
	Aphodius hypocoprus Gistel, 1857 -- invalid
	Aphodius inermis Robinson, 1940 -- invalid
	Aphodius inquinatulus Reitter, 1898 -- invalid
	Aphodius interruptus Mulsant, 1842 -- invalid
	Aphodius inutilis Horn, 1887 -- invalid
	Aphodius inyoensis Van Dyke, 1947 -- invalid
	Aphodius juvenilis Mulsant, 1842 -- invalid
	Aphodius kansanus Wickham, 1905 -- invalid
	Aphodius knausii Fall, 1905 -- invalid
	Aphodius korbeli Roubal, 1935 -- invalid
	Aphodius lecontei Balthasar, 1941 -- invalid
	Aphodius libyanus Balthasar, 1961 -- invalid
	Aphodius limitinus Schmidt, 1922 -- invalid
	Aphodius lineatus Dalla Torre, 1879 -- invalid
	Aphodius lividipennis Waterhouse, 1875 -- invalid
	Aphodius longitarsis Fall, 1907 -- invalid
	Aphodius lopezromeui B�guena, 1927 -- invalid
	Aphodius lucens Stephens, 1830 -- invalid
	Aphodius lunatus Mulsant, 1842 -- invalid
	Aphodius lunulatus Eichler, 1922 -- invalid
	Aphodius maculicollis Montrouzier, 1860 -- invalid
	Aphodius maculipennis Melsheimer, 1845 -- invalid
	Aphodius magnopunctatus Hinton, 1934 -- invalid
	Aphodius marginatus LeConte, 1878 -- invalid
	Aphodius matsuzawai Yawata, 1943 -- invalid
	Aphodius matusitai Nakane, 1961 -- invalid
	Aphodius mcdonaldi Robinson, 1946 -- invalid
	Aphodius melancholicus Dalla Torre, 1879 -- invalid
	Aphodius melanopus Stephens, 1830 -- invalid
	Aphodius melbournicus Balthasar, 1942 -- invalid
	Aphodius metallicus Haldeman, 1848 -- invalid
	Aphodius michiliensis Deloya, 1991 -- invalid
	Aphodius moclipsi Hatch, 1971 -- invalid
	Aphodius mulsanti d'Orbigny, 1896 -- invalid
	Aphodius mundus Reitter, 1892 -- invalid
	Aphodius muticus Stephens, 1830 -- invalid
	Aphodius nama Kolbe, 1908 -- invalid
	Aphodius nanus Horn, 1887 -- invalid
	Aphodius niger Creutzer, 1799 -- invalid
	Aphodius niger Cockerell, 1888 -- invalid
	Aphodius nigritulus Boheman, 1857 -- invalid
	Aphodius nodifrons Randall, 1838 -- invalid
	Aphodius nubiloides Strand and Gulbis, 1957 -- invalid
	Aphodius obliquus Mulsant, 1842 -- invalid
	Aphodius oblongus Say, 1823 -- invalid
	Aphodius obscurus Eichler, 1922 -- invalid
	Aphodius obsoletus Fabricius, 1801 -- invalid
	Aphodius occidentalis Horn, 1870 -- invalid
	Aphodius omiltemius Bates, 1889 -- invalid
	Aphodius ophthalmicus Mulsant, 1842 -- invalid
	Aphodius orophilus de Charpentier, 1825 -- invalid
	Aphodius paivanus Wollaston, 1867 -- invalid
	Aphodius parcepunctatus Mulsant, 1842 -- invalid
	Aphodius pauper Mulsant, 1842 -- invalid
	Aphodius pennsylvanicus Robinson, 1940 -- invalid
	Aphodius pensvallensis Melsheimer, 1845 -- invalid
	Aphodius perpolitus Gordon, 1973 -- invalid
	Aphodius pervicax Balthasar, 1942 -- invalid
	Aphodius phaeopterus LeConte, 1878 -- invalid
	Aphodius pirinensis Balthasar, 1946 -- invalid
	Aphodius politus Horn, 1870 -- invalid
	Aphodius pseudonubilus Strand and Gulbis, 1957 -- invalid
	Aphodius pugil Balthasar, 1946 -- invalid
	Aphodius pulcheroides Strand and Gulbis, 1957 -- invalid
	Aphodius pullmani Saylor, 1940 -- invalid
	Aphodius pumilus Horn, 1887 -- invalid
	Aphodius ragusanus Reitter, 1906 -- invalid
	Aphodius rainieri Hatch, 1971 -- invalid
	Aphodius rapax Faldermann, 1835 -- invalid
	Aphodius rehni Robinson, 1938 -- invalid
	Aphodius restrictus Mulsant, 1842 -- invalid
	Aphodius retusus Waltl, 1835 -- invalid
	Aphodius richardsonae Hatch, 1971 -- invalid
	Aphodius rubidus LeConte, 1857 -- invalid
	Aphodius rufoplagiatus Reitter, 1892 -- invalid
	Aphodius rufotestaceus Dalla Torre, 1879 -- invalid
	Aphodius rugifrons Horn, 1871 -- invalid
	Aphodius rugosulus Mulsant, 1842 -- invalid
	Aphodius russelli Hatch, 1971 -- invalid
	Aphodius sagittarius Harold, 1860 -- invalid
	Aphodius schuhi Gordon, 1975 -- invalid
	Aphodius scutellaris Mulsant, 1842 -- invalid
	Aphodius scutellaris Roth, 1851 -- invalid
	Aphodius seinheili Harold, 1869 -- invalid
	Aphodius sellatus Mannerheim, 1852 -- invalid
	Aphodius semicolor Reitter, 1892 -- invalid
	Aphodius semilunus Mulsant, 1842 -- invalid
	Aphodius semipellitus Solsky, 1876 -- invalid
	Aphodius semiruber Motschulsky, 1860 -- invalid
	Aphodius sequens Walker, 1858 -- invalid
	Aphodius sheldoni Barrett, 1932 -- invalid
	Aphodius signatus Schmidt, 1916 -- invalid
	Aphodius smithi Brown, 1930 -- invalid
	Aphodius snowi Brown, 1929 -- invalid
	Aphodius socialis Brown, 1928 -- invalid
	Aphodius solskyi Harold, 1871 -- invalid
	Aphodius spermophili Gordon, 1973 -- invalid
	Aphodius spilopterus Germar, 1848 -- invalid
	Aphodius spretus Haldeman, 1848 -- invalid
	Aphodius steinheili Harold, 1869 -- invalid
	Aphodius stercoris Gistel, 1857 -- invalid
	Aphodius striatulus Eichler, 1922 -- invalid
	Aphodius striolatus Gebler, 1832 -- invalid
	Aphodius subcarinatus Robinson, 1946 -- invalid
	Aphodius subcinctus Mulsant, 1842 -- invalid
	Aphodius subconfluens Strand and Gulbis, 1957 -- invalid
	Aphodius suspectus Mann, 1911 -- invalid
	Aphodius suturalis Faldermann, 1835 -- invalid
	Aphodius syriacus Mulsant and Rey, 1870 -- invalid
	Aphodius taeniatus Schilsky, 1888 -- invalid
	Aphodius tarsalis Schmidt, 1907 -- invalid
	Aphodius thoracicus Dalla Torre, 1879 -- invalid
	Aphodius tjanshanicus Balthasar, 1956 -- invalid
	Aphodius torpidus Horn, 1870 -- invalid
	Aphodius transversus Robinson, 1940 -- invalid
	Aphodius trifasciatus Strand and Gulbis, 1957 -- invalid
	Aphodius troglodytes Hubbard, 1894 -- invalid -- Aphodius tortoise commensal scarab
	Aphodius tshuensis Protsenko, 1979 -- invalid
	Aphodius tuberifrons Fairmaire, 1903 -- invalid
	Aphodius unicolor Dalla Torre, 1879 -- invalid
	Aphodius ursinus Motschulsky, 1845 -- invalid
	Aphodius utahensis Robinson, 1938 -- invalid
	Aphodius validus Horn, 1870 -- invalid
	Aphodius vestiarius Horn, 1870 -- invalid
	Aphodius vitiosus Reitter, 1892 -- invalid
	Aphodius vitta Motschulsky, 1866 -- invalid
	Aphodius weberi Reitter, 1906 -- invalid
	Aphodius wolcotti Blatchley, 1910 -- invalid
	Aphodius yukonensis Robinson, 1948 -- invalid
	Aphodius zichyi Csiki, 1901 -- invalid
	Aphodius zighanicus Eichler, 1922 -- invalid
	Aphodosugrames Endr�di, 1964 -- invalid
	Aphonides Rivers, 1889 -- invalid
	Aphonus LeConte, 1856 -- valid
	Aphonus aterrimus Casey, 1915 -- invalid
	Aphonus brevicruris Cartwright, 1944 -- valid
	Aphonus castaneus (Melsheimer, 1845) -- valid
	Aphonus congestus Casey, 1915 -- invalid
	Aphonus cubiformis Casey, 1915 -- invalid
	Aphonus densicauda Casey, 1915 -- valid
	Aphonus elongatus Casey, 1915 -- invalid
	Aphonus frater LeConte, 1856 -- invalid
	Aphonus hydropicus LeConte, 1856 -- invalid
	Aphonus ingens Casey, 1924 -- invalid
	Aphonus modulatus Casey, 1915 -- invalid
	Aphonus politus Casey, 1915 -- invalid
	Aphonus saginatus Casey, 1915 -- invalid
	Aphonus scutellaris Casey, 1924 -- invalid
	Aphonus texanus Gill and Howden, 1985 -- valid
	Aphonus trapezicollis Casey, 1915 -- invalid
	Aphonus tridentatus (Say, 1823) -- valid
	Aphonus variolosus (LeConte, 1847) -- valid
	Aphotaenius Cartwright, 1952 -- valid
	Aphotaenius carolinus (Van Dyke, 1928) -- valid
	Aphotaenius howdeni Cartwright, 1963 -- valid
	Apleopertha Semenov, 1903 -- invalid
	Aponelides Kuwert, 1896 -- invalid
	Archimedius Kirby, 1827 -- invalid
	Archophileurus Kolbe, 1910 -- valid
	Archophileurus brevis Casey, 1915 -- invalid
	Archophileurus bullatus Casey, 1915 -- invalid
	Archophileurus cribrosus (LeConte, 1854) -- valid
	Archophileurus longulus Casey, 1915 -- invalid
	Asiopertha Medvedev, 1949 -- invalid
	Aspidolella Prell, 1936 -- invalid
	Aspidolites H�hne, 1922 -- invalid
	Asseroder Maunders, 1848 -- invalid
	Ataeniomorphus Balthasar, 1953 -- invalid
	Ataeniopsis Petrovitz, 1973 -- valid
	Ataeniopsis duncani (Cartwright, 1974) -- valid
	Ataeniopsis edistoi (Cartwright, 1974) -- valid
	Ataeniopsis figurator (Harold, 1874) -- valid
	Ataeniopsis parkeri (Cartwright, 1974) -- valid
	Ataeniopsis rugopygus (Cartwright, 1974) -- valid
	Ataeniopsis saxatilis (Cartwright, 1944) -- valid
	Ataenius Harold, 1867 -- valid
	Ataenius abditus (Haldeman, 1848) -- valid
	Ataenius aequalis Harold, 1880 -- valid
	Ataenius alegrus Balthasar, 1947 -- invalid
	Ataenius alternatus (Melsheimer, 1845) -- valid
	Ataenius anticus Fall, 1930 -- invalid
	Ataenius apicalis Hinton, 1937 -- valid
	Ataenius arator Harold, 1869 -- invalid
	Ataenius barberi Cartwright, 1974 -- valid
	Ataenius benjaminbanderai Islas, 1955 -- invalid
	Ataenius boucomontii Paulian, 1937 -- invalid
	Ataenius braunarum Dury, 1902 -- invalid
	Ataenius brevicollis (Wollaston, 1854) -- valid
	Ataenius brevinotus Chapin, 1940 -- invalid
	Ataenius brevis Fall, 1930 -- valid
	Ataenius californicus Horn, 1887 -- valid
	Ataenius carinator Harold, 1874 -- valid
	Ataenius castaniellus Bates, 1887 -- valid
	Ataenius cognatus (LeConte, 1858) -- valid
	Ataenius confertus Fall, 1909 -- valid
	Ataenius consors Fall, 1930 -- invalid
	Ataenius crenulatus Schmidt, 1910 -- valid
	Ataenius cribratus Van Dyke, 1928 -- invalid
	Ataenius cribrithorax Bates, 1887 -- valid
	Ataenius cylindrus Horn, 1871 -- valid
	Ataenius darlingtoni Hinton, 1937 -- invalid
	Ataenius degallieri Chalumeau, 1990 -- invalid
	Ataenius desertus Horn, 1871 -- valid
	Ataenius duplopunctatus Lea, 1923 -- invalid
	Ataenius ecruensis Stebnicka and Lago, 2005 -- valid
	Ataenius edwardsi Chapin, 1940 -- invalid
	Ataenius erratus Fall, 1930 -- valid
	Ataenius euglyptus Bates, 1887 -- invalid
	Ataenius exiguus Brown, 1932 -- valid
	Ataenius falli Hinton, 1934 -- invalid
	Ataenius fattigi Cartwright, 1948 -- valid
	Ataenius fleutiauxi Paulian, 1947 -- invalid
	Ataenius floridanus Brown, 1930 -- invalid
	Ataenius frankenbergeri Balthasar, 1938 -- invalid
	Ataenius glaseri Cartwright, 1974 -- valid
	Ataenius gracilis (Melsheimer, 1845) -- valid
	Ataenius granchaocensis Balthasar, 1939 -- invalid
	Ataenius havanensis Balthasar, 1938 -- invalid
	Ataenius heinekeni (Wollaston, 1854) -- valid
	Ataenius hesperius Cartwright, 1974 -- valid
	Ataenius heyrovskyi Balthasar, 1960 -- invalid
	Ataenius hieronymi Bates, 1887 -- invalid
	Ataenius hirsutus Horn, 1871 -- valid
	Ataenius histrionicus Balthasar, 1947 -- invalid
	Ataenius hoguei Cartwright and Spangler, 1981 -- invalid
	Ataenius horni Harold, 1874 -- invalid
	Ataenius imbricatus (Melsheimer, 1845) -- valid
	Ataenius inops Horn, 1887 -- invalid
	Ataenius inquisitus Horn, 1887 -- valid
	Ataenius insculptus Horn, 1887 -- valid
	Ataenius insulicola Chapin, 1940 -- invalid
	Ataenius integer Harold, 1868 -- invalid
	Ataenius jalapensis Bates, 1887 -- invalid
	Ataenius kelatianus Balthasar, 1965 -- invalid
	Ataenius laeviventris Horn, 1887 -- invalid
	Ataenius languidus Schmidt, 1911 -- valid
	Ataenius lecontei Harold, 1874 -- invalid
	Ataenius lherminieri Paulian, 1947 -- invalid
	Ataenius linelli Cartwright, 1944 -- invalid
	Ataenius liogaster Bates, 1887 -- valid
	Ataenius lobatus Horn, 1871 -- valid
	Ataenius ludovicianus Fall, 1930 -- invalid
	Ataenius lusitanicus Paulian, 1979 -- invalid
	Ataenius miamii Cartwright, 1934 -- valid
	Ataenius narialis Petrovitz, 1973 -- invalid
	Ataenius nitidulus Nomura, 1943 -- invalid
	Ataenius nunenmacheri Cartwright, 1974 -- invalid
	Ataenius oblongus Horn, 1871 -- invalid
	Ataenius oklahomensis Brown, 1930 -- valid
	Ataenius opatrinus Harold, 1867 -- valid
	Ataenius orbicularis Schmidt, 1914 -- invalid
	Ataenius ovatulus Horn, 1871 -- valid
	Ataenius paracognatus Balthasar, 1961 -- invalid
	Ataenius paraperforatus Deloya and Ib��ez-Bernal, 2000 -- invalid
	Ataenius peregrinator Harold, 1877 -- valid
	Ataenius picinus Harold, 1867 -- valid
	Ataenius platensis (Blanchard, 1847) -- valid
	Ataenius pseudohirsutus Cartwright, 1974 -- invalid
	Ataenius punctifrons Cartwright, 1974 -- invalid
	Ataenius queirosii Paulian, 1934 -- invalid
	Ataenius rickardsi Hinton, 1938 -- invalid
	Ataenius robustus Horn, 1871 -- valid
	Ataenius rudellus Fall, 1930 -- invalid
	Ataenius rugiceps Dury, 1902 -- invalid
	Ataenius sabinoi Cartwright, 1974 -- invalid
	Ataenius salutator Fall, 1930 -- invalid
	Ataenius scabrelloides Petrovitz, 1962 -- valid
	Ataenius scabrellus Schmidt, 1909 -- valid
	Ataenius sciurus Cartwright, 1974 -- invalid
	Ataenius sculptor Harold, 1868 -- valid
	Ataenius setiger Bates, 1887 -- valid
	Ataenius solitarius Blatchley, 1928 -- invalid
	Ataenius sordidus Harold, 1869 -- invalid
	Ataenius spretulus (Haldeman, 1848) -- valid
	Ataenius stephani Cartwright, 1974 -- valid
	Ataenius strigatus (Say, 1823) -- valid
	Ataenius stroheckeri Cartwright, 1974 -- invalid
	Ataenius superficialis Cartwright, 1974 -- invalid -- Big Pine Key ataenius dung beetle
	Ataenius texanus Harold, 1874 -- valid
	Ataenius titschacki Balthasar, 1941 -- invalid
	Ataenius utahensis Cartwright, 1974 -- valid
	Ataenius vandykei Cartwright, 1974 -- valid
	Ataenius vexator Harold, 1869 -- invalid
	Ataenius vincentiae Arrow, 1903 -- invalid
	Ataenius waltherhorni Balthasar, 1938 -- valid
	Ataenius wenzelii Horn, 1887 -- valid
	Ataenius woodruffi Cartwright, 1974 -- invalid -- Woodruff's ataenius dung beetle
	Ateuchini Perty, 1830 -- valid
	Ateuchus Weber, 1801 -- valid
	Ateuchus capistratus Fabricius, 1801 -- invalid
	Ateuchus floridensis G�nier, 2000 -- valid
	Ateuchus histeroides Weber, 1801 -- valid
	Ateuchus illaesum (Harold, 1868) -- valid
	Ateuchus lecontei (Harold, 1868) -- valid
	Ateuchus obsoletus Say, 1823 -- invalid
	Ateuchus rodriguezi (Preudhomme de Borre, 1886) -- valid
	Ateuchus texanus (Robinson, 1948) -- valid
	Athyreini Lynch Arrib�lzaga, 1878 -- valid
	Auperia Jacquelin du Val, 1857 -- invalid
	Auperia denominata Chevrolat, 1864 -- invalid
	Auperia rhyticephalus Chevrolat, 1864 -- invalid
	Auperia sulcatula Chevrolat, 1864 -- invalid
	Australaphodius Balthasar, 1942 -- valid
	Australaphodius frenchi (Blackburn, 1892) -- valid
	Aztecalius Casey, 1915 -- invalid
	Ballucus Gordon and Skelley, 2007 -- valid
	Ballucus barri (Gordon, 2006) -- valid
	Ballucus bruneaui Gordon and Skelley, 2007 -- valid
	Ballucus obtusus (LeConte, 1878) -- valid
	Ballucus oregonus Gordon and Skelley, 2007 -- valid
	Barotheus Bates, 1891 -- invalid
	Barymorpha Gu�rin-M�neville, 1843 -- invalid
	Benedictia Sanderson, 1939 -- invalid
	Blackburneus Schmidt, 1913 -- valid
	Blackburneus aegrotus (Horn, 1870) -- invalid
	Blackburneus cavidomus (Brown, 1927) -- invalid
	Blackburneus cynomysi (Brown, 1927) -- invalid
	Blackburneus fordi (Gordon, 1974) -- invalid
	Blackburneus geomysi (Cartwright, 1939) -- invalid
	Blackburneus lentus (Horn, 1870) -- invalid
	Blackburneus rubeolus (Palisot de Beauvois, 1809) -- invalid
	Blackburneus saylori (Hinton, 1934) -- invalid
	Blackburneus stercorosus (Melsheimer, 1844) -- valid
	Blackburneus tenuistriatus (Horn, 1887) -- invalid
	Blackburneus troglodytes (Hubbard, 1894) -- invalid
	Blaesiina Schoch, 1895 -- valid
	Blitopertha Reitter, 1903 -- invalid
	Blitopertha flavipennis Reitter, 1903 -- invalid
	Bolbelasmus Boucomont, 1911 -- valid
	Bolbelasmus arcuatus (Bates, 1887) -- valid
	Bolbelasmus bajaensis (Howden, 1964) -- valid
	Bolbelasmus hornii (Rivers, 1886) -- valid
	Bolbelasmus minor (Linell, 1896) -- valid
	Bolbelasmus variabilis Howden, 1964 -- valid
	Bolboceras furcicollis Laporte, 1840 -- invalid
	Bolboceras schaefferi Boucomont, 1911 -- invalid
	Bolbocerastes Cartwright, 1953 -- valid
	Bolbocerastes imperialis Cartwright, 1953 -- valid
	Bolbocerastes imperialis imperialis Cartwright, 1953 -- valid
	Bolbocerastes imperialis kansanus Cartwright, 1953 -- valid
	Bolbocerastes peninsularis (Schaeffer, 1906) -- valid
	Bolbocerastes regalis Cartwright, 1953 -- valid
	Bolbocerastes serratus (LeConte, 1854) -- valid
	Bolboceratinae Mulsant, 1842 -- valid
	Bolboceratini Mulsant, 1842 -- valid
	Bolbocerosoma Schaeffer, 1906 -- valid
	Bolbocerosoma (Bolbocerosoma) Schaeffer, 1906 -- valid
	Bolbocerosoma biplagiatum Dawson and McColloch, 1924 -- valid
	Bolbocerosoma bruneri Dawson and McColloch, 1924 -- valid
	Bolbocerosoma cartwrighti Howden, 1955 -- valid
	Bolbocerosoma confusum Brown, 1928 -- valid
	Bolbocerosoma elongatum Howden, 1955 -- valid
	Bolbocerosoma farctum (Fabricius, 1775) -- valid
	Bolbocerosoma hamatum Brown, 1929 -- valid
	Bolbocerosoma lepidissimum Brown, 1928 -- valid
	Bolbocerosoma mexicanum Howden, 2005 -- valid
	Bolbocerosoma pusillum Dawson and McColloch, 1924 -- valid
	Bolbocerosoma pusillum pusillum Dawson and McColloch, 1924 -- valid
	Bolbocerosoma pusillum townesi Howden, 1955 -- valid
	Bolbocerosoma quadricornum Robinson, 1941 -- valid
	Bolbocerosoma ritcheri Howden, 1955 -- valid
	Bolbocerosoma tumefactum (Palisot de Beauvois, 1809) -- valid
	Bolborhombus Cartwright, 1953 -- valid
	Bolborhombus angulus (Robinson, 1947) -- valid
	Bolborhombus magnus Howden, 1964 -- valid
	Bolborhombus parvulus Cartwright, 1953 -- valid
	Bolborhombus sallaei (Bates, 1887) -- valid
	Bothynus morio LeConte, 1847 -- invalid
	Bothynus obsoletus LeConte, 1847 -- invalid
	Boucardius Petrovitz, 1967 -- invalid
	Brachycopris Haldeman, 1846 -- invalid
	Bradycinetulus Cockerell, 1906 -- valid
	Bradycinetulus ferrugineus (Palisot de Beauvois, 1809) -- valid
	Bradycinetulus fossatus (Haldeman, 1853) -- valid
	Bradycinetulus rex Cartwright, 1953 -- valid
	Bradycinetus Horn, 1871 -- invalid
	Bradycinetus carinatus Schaeffer, 1906 -- invalid
	Brancoataenius Paulian, 1979 -- invalid
	Byrsopolis arizonae Ohaus, 1912 -- invalid
	Byrsopolis lanigera Bates, 1888 -- invalid
	Caelius Lewis, 1895 -- valid
	Caelius browni (Saylor, 1934) -- valid
	Caelius humeralis (Brown, 1931) -- valid
	Caelius montana (Brown, 1931) -- valid
	Caelius rufescens (Horn, 1887) -- valid
	Calamosternus Motschulsky, 1859 -- valid
	Calamosternus granarius (Linnaeus, 1767) -- valid
	Calidas Kuwert, 1896 -- invalid
	Caligodorus Gordon and Skelley, 2007 -- valid
	Caligodorus opacus (LeConte, 1872) -- valid
	Caligodorus tuberosus (Barrett, 1932) -- valid
	Caligodorus vandykei (Barrett, 1931) -- valid
	Callistethus Blanchard, 1851 -- invalid
	Calomacraspis Bates, 1888 -- valid
	Calomacraspis splendens (Burmeister, 1844) -- valid
	Calopopillia Kolbe, 1894 -- invalid
	Camptorhina Kirby, 1837 -- invalid
	Canthidium Erichson, 1847 -- valid
	Canthidium (Eucanthidium) Mart�nez and Halffter, 1986 -- valid
	Canthidium laetum Harold, 1867 -- valid
	Canthidium macclevei Kohlmann and Sol�s, 2006 -- valid
	Canthidium pseudopuncticolle Sol�s and Kohlmann, 2004 -- valid
	Canthomoechus Pereira and Mart�nez, 1960 -- invalid
	Canthon Hoffmannsegg, 1817 -- valid
	Canthon (Boreocanthon) Halffter, 1958 -- valid
	Canthon (Canthon) Hoffmannsegg, 1817 -- valid
	Canthon (Glaphyrocanthon) Mart�nez, 1948 -- valid
	Canthon abrasus LeConte, 1859 -- invalid
	Canthon amethystinum Harold, 1863 -- invalid
	Canthon antiquus Pierce, 1946 -- invalid
	Canthon balsensis Mart�nez and Halffter, 1972 -- invalid
	Canthon bisignatus Balthasar, 1939 -- valid
	Canthon chalcites (Haldeman, 1843) -- valid
	Canthon chevrolati Harold, 1868 -- invalid
	Canthon coahuilensis Howden, 1966 -- valid
	Canthon corvinus Harold, 1868 -- invalid
	Canthon cyanellus LeConte, 1859 -- valid
	Canthon depressipennis LeConte, 1859 -- valid
	Canthon ebenus (Say, 1823) -- valid
	Canthon femoralis (Chevrolat, 1834) -- valid
	Canthon floridanus Brown, 1946 -- invalid
	Canthon forreri Bates, 1887 -- valid
	Canthon gagatinus Harold, 1863 -- invalid
	Canthon humectus (Say, 1832) -- valid
	Canthon humectus alvarengai Halffter, 1961 -- valid
	Canthon humectus blumei Halffter and Halffter, 2003 -- valid
	Canthon humectus hidalgoensis Bates, 1887 -- valid
	Canthon humectus humectus (Say, 1832) -- valid
	Canthon humeralis Horn, 1870 -- invalid
	Canthon imitator Brown, 1946 -- valid
	Canthon indigaceus LeConte, 1866 -- valid
	Canthon integricollis Schaeffer, 1915 -- invalid
	Canthon laevis Bates, 1887 -- invalid
	Canthon lecontei Harold, 1868 -- valid
	Canthon melanus Robinson, 1948 -- valid
	Canthon metallicus Sturm, 1843 -- invalid
	Canthon militaris Horn, 1870 -- invalid
	Canthon minor Sturm, 1843 -- invalid
	Canthon mixtus Robinson, 1948 -- valid
	Canthon nyctelius Bates, 1887 -- invalid
	Canthon obliquus Horn, 1894 -- valid
	Canthon pilularius (Linnaeus, 1758) -- valid
	Canthon praticola LeConte, 1859 -- valid
	Canthon probus (Germar, 1824) -- valid
	Canthon puncticollis LeConte, 1866 -- valid
	Canthon simplex LeConte, 1857 -- valid
	Canthon vetustus Pierce, 1946 -- invalid
	Canthon vigilans LeConte, 1858 -- valid
	Canthon viridescens Horn, 1870 -- invalid
	Canthon viridis (Palisot de Beauvois, 1805) -- valid
	Canthonini van Lansberge, 1875 -- valid
	Cassius Kuwert, 1891 -- invalid
	Castapopillia Benderitter, 1922 -- invalid
	Centrochilus Krikken, 1976 -- valid
	Centrochilus howdeni Krikken, 1976 -- valid
	Cephalocyclus Dellacasa, Gordon and Dellacasa, 1998 -- valid
	Cephalocyclus bordati Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Cephalocyclus carmenae Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Cephalocyclus durangoensis (Bates, 1887) -- valid
	Cephalocyclus fuliginosus (Harold, 1863) -- valid
	Cephalocyclus gravidus (Harold, 1863) -- valid
	Cephalocyclus hogei (Bates, 1887) -- valid
	Cephalocyclus howdenorum Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Cephalocyclus lagoi Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Cephalocyclus luteolus (Horn, 1887) -- valid
	Cephalocyclus mexicanus (Harold, 1862) -- valid
	Cephalocyclus potosinus Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Cephalocyclus pullatus (Schmidt, 1913) -- valid
	Cephalocyclus rockefelleri Dellacasa, Dellacasa and Gordon, 2007 -- valid
	Ceratocanthidae Mart�nez, 1968 -- invalid
	Ceratocanthinae Mart�nez, 1968 -- valid
	Ceratocanthini Mart�nez, 1968 -- valid
	Ceratocanthus White, 1842 -- valid
	Ceratocanthus aeneus (MacLeay, 1819) -- valid
	Ceratophyus Fischer von Waldheim, 1823 -- valid
	Ceratophyus gopherinus Cartwright, 1966 -- valid
	Ceratotrupes Jekel, 1865 -- valid
	Ceratotrupes bolivari Halffter and Mart�nez, 1962 -- valid
	Ceruchus MacLeay, 1819 -- valid
	Ceruchus piceus (Weber, 1801) -- valid
	Ceruchus punctatus LeConte, 1869 -- valid
	Ceruchus striatus LeConte, 1859 -- valid
	Ceruchus virginiensis Casey, 1914 -- invalid
	Cetonia antennata Gory and Percheron, 1833 -- invalid
	Cetonia atomaria Fabricius, 1801 -- invalid
	Cetonia barbata Say, 1824 -- invalid
	Cetonia brunnea Gory and Percheron, 1833 -- invalid
	Cetonia childrenii Gory and Percheron, 1833 -- invalid
	Cetonia ferrugata Gory and Percheron, 1833 -- invalid
	Cetonia fictilis Newman, 1838 -- invalid
	Cetonia mandarina Lichtenstein, 1796 -- invalid
	Cetonia marilandica Fr�lich, 1792 -- invalid
	Cetonia montesuma Gory and Percheron, 1833 -- invalid
	Cetonia pubera Gyllenhal, 1817 -- invalid
	Cetonia reichii Gory and Percheron, 1833 -- invalid
	Cetonia thelasco Gory and Percheron, 1833 -- invalid
	Cetoniina Leach, 1815 -- valid
	Cetoniinae Leach, 1815 -- valid
	Cetoniini Leach, 1815 -- valid
	Chaetocanthinae Scholtz in Scholtz, D'hotman, Evans and Nel, 1988 -- valid
	Chalcoderus Erichson, 1848 -- invalid
	Chalcoplethis Burmeister, 1844 -- invalid
	Chalepus MacLeay, 1819 -- invalid
	Chalepus leucophthalma Fischer von Waldheim, 1823 -- invalid
	Chalepus obsoletus LeConte, 1854 -- invalid
	Chalepus trachypygus Burmeister, 1847 -- invalid
	Chaunocolus Saylor, 1937 -- valid
	Chaunocolus cornutus Saylor, 1937 -- valid
	Chesas Burmeister, 1876 -- invalid
	Chilothorax Motschulsky, 1859 -- valid
	Chilothorax distinctus (M�ller, 1776) -- valid
	Chlorixanthe Bates, 1889 -- valid
	Chlorixanthe chapini Cartwright, 1939 -- invalid
	Chlorixanthe propinqua (Gory and Percheron, 1833) -- valid
	Chnaunanthus Burmeister, 1844 -- valid
	Chnaunanthus chapini Saylor, 1937 -- valid
	Chnaunanthus discolor Burmeister, 1844 -- valid
	Chnaunanthus flavipennis (Horn, 1867) -- valid
	Chnaunanthus palmeri Horn, 1894 -- invalid
	Choeridium LePeletier and Serville, 1828 -- invalid
	Choeridium punctatus Robinson, 1948 -- invalid
	Chremastodus Solier, 1851 -- invalid
	Chrysina Kirby, 1828 -- valid
	Chrysina adelaida (Hope, 1840) -- valid
	Chrysina beckeri Bates, 1889 -- valid
	Chrysina beyeri (Skinner, 1905) -- valid
	Chrysina brevis (Rothschild and Jordan, 1894) -- valid
	Chrysina erubescens Bates, 1889 -- valid
	Chrysina flohri (Ohaus, 1905) -- valid
	Chrysina gloriosa (LeConte, 1854) -- valid
	Chrysina lecontei (Horn, 1882) -- valid
	Chrysina plusiotina (Ohaus, 1912) -- valid
	Chrysina psittacina (Sturm, 1843) -- valid
	Chrysina woodii (Horn, 1885) -- valid
	Cinacanthus Schmidt, 1913 -- valid
	Cinacanthus coquilletti (Linell, 1896) -- valid
	Cinacanthus crenicollis (Fall, 1932) -- valid
	Cinacanthus crescentus Gordon and Skelley, 2007 -- valid
	Cinacanthus hirsutus (Brown, 1928) -- valid
	Cinacanthus militaris (LeConte, 1858) -- valid
	Cinacanthus ulkei (Gordon, 2006) -- valid
	Cineretis Krikken, 1984 -- invalid
	Clemora Saylor, 1942 -- invalid
	Cloeotus prionomus Bates, 1887 -- invalid
	Cnemargus Harold, 1866 -- invalid
	Codocera Eschscholtz, 1821 -- valid
	Codocera gnatho (Fall, 1907) -- valid
	Coeloscelis Reiche, 1841 -- invalid
	Coelotrachelus Schmidt, 1913 -- valid
	Coelotrachelus kuntzeni (Schmidt, 1913) -- valid
	Coelotrachelus rudis (LeConte, 1878) -- valid
	Coelotrachelus symbius Gordon and Howden, 1973 -- valid
	Coenonycha Horn, 1876 -- valid
	Coenonycha acuta Cazier, 1943 -- valid
	Coenonycha ampla Cazier, 1943 -- valid
	Coenonycha barri Cazier, 1943 -- valid
	Coenonycha bowlesi Cazier, 1943 -- valid
	Coenonycha clementina Casey, 1909 -- valid -- San Clemente Island coenonycha beetle
	Coenonycha clypeata McClay, 1943 -- valid
	Coenonycha crispata McClay, 1943 -- valid
	Coenonycha dimorpha Evans, 1986 -- valid
	Coenonycha fuga Cazier, 1943 -- valid
	Coenonycha fulva McClay, 1943 -- valid
	Coenonycha fusca McClay, 1943 -- valid
	Coenonycha globosa McClay, 1943 -- valid
	Coenonycha hageni Cazier, 1943 -- valid
	Coenonycha lurida Cazier, 1943 -- valid
	Coenonycha mediata Cazier, 1943 -- valid
	Coenonycha ochreata Evans, 1986 -- valid
	Coenonycha ovatis McClay, 1943 -- valid
	Coenonycha ovipennis Horn, 1876 -- valid
	Coenonycha pallida Cazier, 1943 -- valid
	Coenonycha parvula Fall, 1901 -- valid
	Coenonycha pascuensis Potts, 1945 -- valid
	Coenonycha purshiae Cazier, 1943 -- valid
	Coenonycha pygmaea Smith, 1986 -- valid
	Coenonycha rotundata (LeConte, 1856) -- valid
	Coenonycha rubida McClay, 1943 -- valid
	Coenonycha rufobrunnea Howden, 1969 -- invalid
	Coenonycha santacruzae Evans, 1986 -- valid
	Coenonycha saylori Cazier, 1943 -- valid
	Coenonycha scotti McClay, 1943 -- valid
	Coenonycha sleeperi Evans, 1988 -- valid
	Coenonycha socialis Horn, 1876 -- valid
	Coenonycha stohleri Saylor, 1935 -- valid
	Coenonycha testacea Cazier, 1937 -- valid
	Coenonycha tingi Cazier, 1937 -- valid
	Coenonycha utahensis McClay, 1943 -- valid
	Colobopterus Mulsant, 1842 -- valid
	Colobopterus erraticus (Linnaeus, 1758) -- valid
	Colobopterus fumigatus Mulsant, 1842 -- invalid
	Colobopterus nebulosus Mulsant, 1842 -- invalid
	Colobopterus submaculatus Mulsant, 1842 -- invalid
	Colporhina Curtis, 1844 -- invalid
	Coprini Leach, 1815 -- valid
	Copris Geoffroy, 1762 -- valid
	Copris anaglypticus Say, 1823 -- invalid
	Copris antilope Fabricius, 1798 -- invalid
	Copris arizonensis Schaeffer, 1906 -- valid
	Copris cartwrighti Robinson, 1941 -- invalid
	Copris clavicornis Matthews and Halffter, 1959 -- invalid
	Copris corniger Geoffroy, 1785 -- invalid
	Copris fricator (Fabricius, 1787) -- valid
	Copris gopheri Hubbard, 1894 -- valid -- Copris tortoise commensal scarab
	Copris hastator Fabricius, 1798 -- invalid
	Copris howdeni Matthews and Halffter, 1959 -- valid
	Copris incertus Say, 1835 -- valid
	Copris inemarginatus Blatchley, 1918 -- valid
	Copris klugi sierrensis Matthews, 1962 -- valid
	Copris laeviceps Harold, 1862 -- valid
	Copris latebrosus Fabricius, 1801 -- invalid
	Copris lecontei Matthews, 1962 -- valid
	Copris lecontei lecontei Matthews, 1962 -- valid
	Copris lugubris Boheman, 1858 -- valid
	Copris macclevei Warner, 1990 -- valid
	Copris martinezi Matthews and Halffter, 1968 -- valid
	Copris metallicus Fabricius, 1798 -- invalid
	Copris minutus (Drury, 1773) -- valid
	Copris moechus LeConte, 1854 -- valid
	Copris obtectus Palisot de Beauvois, 1805 -- invalid
	Copris procidua Say, 1835 -- invalid
	Copris reflexus Panzer, 1794 -- invalid
	Copris remotus LeConte, 1866 -- valid
	Copris remotus dicyrtus Matthews and Halffter, 1959 -- valid
	Copris remotus remotus LeConte, 1866 -- valid
	Coprobius Latreille, 1829 -- invalid
	Coprobius obtusidens Ziegler, 1844 -- invalid
	Coprocanthon Mart�nez, 1948 -- invalid
	Coprophanaeus D'Olsoufieff, 1924 -- valid
	Coprophanaeus (Coprophanaeus) D'Olsoufieff, 1924 -- valid
	Coprophanaeus pluto noguerai Arnaud, 2002 -- invalid
	Coscinocephalus Prell, 1936 -- valid
	Coscinocephalus cribrifrons (Schaeffer, 1906) -- valid
	Cotalpa Burmeister, 1844 -- valid
	Cotalpa ashleyae LaRue, 1986 -- valid
	Cotalpa batesi Ohaus, 1915 -- invalid
	Cotalpa conclamara Young, 2002 -- valid
	Cotalpa consobrina Horn, 1871 -- valid
	Cotalpa flavida Horn, 1878 -- valid
	Cotalpa lanigera (Linnaeus, 1758) -- valid
	Cotalpa leonina Fall, 1932 -- invalid
	Cotalpa molaris Casey, 1915 -- invalid
	Cotalpa obesa Casey, 1915 -- invalid
	Cotalpa subcribrata Wickham, 1905 -- valid
	Cotalpa tau Wickham, 1905 -- invalid
	Cotalpa vernicata Casey, 1915 -- invalid
	Cotinis Burmeister, 1842 -- valid
	Cotinis (Cotinis) Burmeister, 1842 -- valid
	Cotinis (Criniflava) Goodrich, 1966 -- valid
	Cotinis abdominalis Casey, 1915 -- invalid
	Cotinis aliena Woodruff, 2008 -- valid
	Cotinis angustula Casey, 1915 -- invalid
	Cotinis antonii (Dug�s, 1878) -- valid
	Cotinis arizonica Casey, 1915 -- invalid
	Cotinis atra Kraatz, 1898 -- invalid
	Cotinis atropurpurea Kraatz, 1898 -- invalid
	Cotinis aurantica Bates, 1889 -- invalid
	Cotinis boylei Goodrich, 1966 -- valid
	Cotinis brunnea Hatch, 1930 -- invalid
	Cotinis cabira Burmeister, 1842 -- invalid
	Cotinis capito Casey, 1915 -- invalid
	Cotinis coahuilae Casey, 1915 -- invalid
	Cotinis commiscens Casey, 1915 -- invalid
	Cotinis cuprascens Bates, 1889 -- invalid
	Cotinis cuprea Kraatz, 1898 -- invalid
	Cotinis debiliceps Casey, 1915 -- invalid
	Cotinis discolor Casey, 1915 -- invalid
	Cotinis impia (Fall, 1905) -- valid
	Cotinis intergenea Bates, 1889 -- invalid
	Cotinis laticornis Bates, 1889 -- valid
	Cotinis latifrons Schoch, 1895 -- invalid
	Cotinis longitarsus Casey, 1915 -- invalid
	Cotinis longula Casey, 1915 -- invalid
	Cotinis malinus Janson, 1889 -- invalid
	Cotinis mutabilis (Gory and Percheron, 1833) -- valid
	Cotinis nigrovariegata Kraatz, 1898 -- invalid
	Cotinis nitida (Linnaeus, 1758) -- valid
	Cotinis obliqua Casey, 1915 -- invalid
	Cotinis orientalis Deloya and Ratcliffe, 1988 -- valid
	Cotinis ornata Casey, 1915 -- invalid
	Cotinis ovicornuta Casey, 1915 -- invalid
	Cotinis parvula Casey, 1915 -- invalid
	Cotinis piciventris Kraatz, 1898 -- invalid
	Cotinis producta Bates, 1889 -- valid
	Cotinis pygidialis Casey, 1915 -- invalid
	Cotinis robusta Bates, 1889 -- invalid
	Cotinis rufipennis Bates, 1889 -- valid
	Cotinis schafraneki Nonfried, 1894 -- invalid
	Cotinis senex Janson, 1880 -- invalid
	Cotinis seriella Casey, 1915 -- invalid
	Cotinis sinitoc Deloya, Iba�ez-Bernal and Nogueira, 2000 -- valid
	Cotinis subcastanca Bates, 1889 -- invalid
	Cotinis texana Casey, 1915 -- invalid
	Cotinis tibialis Casey, 1915 -- invalid
	Cotinis viridicauda Casey, 1915 -- invalid
	Cotinorrhina Schoch, 1895 -- invalid
	Cremastocheilina Burmeister and Schaum, 1841 -- valid
	Cremastocheilini Burmeister and Schaum, 1841 -- valid
	Cremastocheilus Knoch, 1801 -- valid
	Cremastocheilus (Anatrinodia) Casey, 1915 -- valid
	Cremastocheilus (Cremastocheilus) Knoch, 1801 -- valid
	Cremastocheilus (Macropodina) Knoch, 1801 -- valid
	Cremastocheilus (Myrmecotonus) Mann, 1914 -- valid
	Cremastocheilus (Trinodia) Knoch, 1801 -- valid
	Cremastocheilus academicus Krikken, 1982 -- valid
	Cremastocheilus angularis LeConte, 1857 -- valid
	Cremastocheilus areolatus Casey, 1915 -- invalid
	Cremastocheilus armatus Walker, 1866 -- valid
	Cremastocheilus armatus armatus Walker, 1866 -- valid
	Cremastocheilus armatus cribripennis Casey, 1915 -- valid
	Cremastocheilus armatus maritimus Casey, 1915 -- valid
	Cremastocheilus armatus montanus Casey, 1915 -- valid
	Cremastocheilus beameri Cazier, 1940 -- valid
	Cremastocheilus canaliculatus Kirby, 1827 -- valid
	Cremastocheilus castaneae Knoch, 1801 -- valid
	Cremastocheilus castaneae brevisetosus Casey, 1915 -- valid
	Cremastocheilus castaneae castaneae Knoch, 1801 -- valid
	Cremastocheilus castaneae lecontei Westwood, 1874 -- valid
	Cremastocheilus castaneae pocularis Casey, 1915 -- valid
	Cremastocheilus chapini Cazier, 1940 -- valid
	Cremastocheilus cicatricosus Westwood, 1874 -- invalid
	Cremastocheilus congener Casey, 1915 -- valid
	Cremastocheilus constricollis Cazier, 1940 -- valid
	Cremastocheilus crassipes Westwood, 1874 -- invalid
	Cremastocheilus crenicollis Westwood, 1874 -- invalid
	Cremastocheilus crinitus LeConte, 1874 -- valid
	Cremastocheilus crinitus bifoveatus Van Dyke, 1918 -- valid
	Cremastocheilus crinitus crinitus LeConte, 1874 -- valid
	Cremastocheilus crinitus pugetanus Casey, 1915 -- valid
	Cremastocheilus excavatus Cazier, 1940 -- valid
	Cremastocheilus gracilipes Casey, 1915 -- invalid
	Cremastocheilus harrisii Kirby, 1827 -- valid
	Cremastocheilus hentzii Harris, 1827 -- invalid
	Cremastocheilus hirsutus Van Dyke, 1918 -- valid
	Cremastocheilus junior Westwood, 1874 -- invalid
	Cremastocheilus knochii LeConte, 1853 -- valid
	Cremastocheilus lengi Cazier, 1940 -- valid
	Cremastocheilus mentalis Cazier, 1940 -- valid
	Cremastocheilus mexicanus Schaum, 1841 -- valid
	Cremastocheilus nitens LeConte, 1853 -- valid
	Cremastocheilus opaculus Horn, 1894 -- valid
	Cremastocheilus percheronii Westwood, 1874 -- invalid
	Cremastocheilus pilosicollis Horn, 1871 -- invalid
	Cremastocheilus planatus LeConte, 1863 -- valid
	Cremastocheilus planipes Horn, 1885 -- valid
	Cremastocheilus pulverulentus Cazier, 1940 -- valid
	Cremastocheilus puncticollis Cazier, 1940 -- valid
	Cremastocheilus quadratus Fall, 1912 -- valid
	Cremastocheilus quadricollis (Casey, 1915) -- valid
	Cremastocheilus retractus LeConte, 1874 -- valid
	Cremastocheilus retractus incisus Casey, 1915 -- valid
	Cremastocheilus retractus retractus LeConte, 1874 -- valid
	Cremastocheilus robinsoni Cazier, 1940 -- valid
	Cremastocheilus saucius LeConte, 1858 -- valid
	Cremastocheilus sayi Harris, 1827 -- invalid
	Cremastocheilus schaumii LeConte, 1853 -- valid
	Cremastocheilus schaumii schaumii LeConte, 1853 -- valid
	Cremastocheilus schaumii tibialis Casey, 1915 -- valid
	Cremastocheilus setosifrons (Casey, 1915) -- valid
	Cremastocheilus spinifer Horn, 1885 -- valid
	Cremastocheilus squamulosus LeConte, 1858 -- valid
	Cremastocheilus stathamae Cazier, 1961 -- valid
	Cremastocheilus tomentosus Warner, 1985 -- valid
	Cremastocheilus variolosus Kirby, 1826 -- valid
	Cremastocheilus villadae Dug�s, 1879 -- invalid
	Cremastocheilus walshii Westwood, 1874 -- invalid
	Cremastocheilus westwoodi Horn, 1880 -- valid
	Cremastocheilus westwoodi tridens Casey, 1915 -- valid
	Cremastocheilus westwoodi westwoodi Horn, 1880 -- valid
	Cremastocheilus wheeleri LeConte, 1876 -- valid
	Cryptoscatomaseter Gordon and Skelley, 2007 -- valid
	Cryptoscatomaseter acuminatus (Cartwright, 1944) -- valid
	Cryptoscatomaseter brevicollis (LeConte, 1878) -- valid
	Cryptoscatomaseter browni (Hinton, 1934) -- valid
	Cryptoscatomaseter caccabatus (Warner and Skelley, 2006) -- valid
	Cryptoscatomaseter captivus (Cartwright, 1944) -- valid
	Cryptoscatomaseter criddlei (Brown, 1928) -- valid
	Cryptoscatomaseter depressiusculus (Schmidt, 1907) -- valid
	Cryptoscatomaseter durangoi (Gordon and Howden, 1973) -- valid
	Cryptoscatomaseter ejectus Gordon and Skelley, 2007 -- valid
	Cryptoscatomaseter epulus (Gordon and Howden, 1973) -- valid
	Cryptoscatomaseter excavatus (Robinson, 1940) -- valid
	Cryptoscatomaseter gambrinus (Skelley and Gordon, 2002) -- valid
	Cryptoscatomaseter haldemani (Horn, 1887) -- valid
	Cryptoscatomaseter hubbelli (Skelley and Woodruff, 1991) -- valid
	Cryptoscatomaseter incomptus Gordon and Skelley, 2007 -- valid
	Cryptoscatomaseter iowensis (Wickham, 1913) -- valid
	Cryptoscatomaseter magnificens (Robinson, 1940) -- valid
	Cryptoscatomaseter ochreipennis (Horn, 1871) -- valid
	Cryptoscatomaseter oklahomensis (Brown, 1928) -- valid
	Cryptoscatomaseter paulseni (Skelley, 2006) -- valid
	Cryptoscatomaseter pholetus (Skelley and Woodruff, 1991) -- valid
	Cryptoscatomaseter platypleurus (Skelley and Woodruff, 1991) -- valid
	Cryptoscatomaseter punctissimus (Brown, 1928) -- valid
	Cryptoscatomaseter rossi (Cartwright, 1944) -- valid
	Cryptoscatomaseter salsburyi (Gordon, 2006) -- valid
	Cryptoscatomaseter talpoidesi (Brown, 1928) -- valid
	Cryptoscatomaseter umbricollis (Fall, 1907) -- valid
	Cryptoscatomaseter utopensis (Warner and Skelley, 2006) -- valid
	Cryptoscatomaseter yavapai (Robinson, 1946) -- valid
	Cucochodaeus Paulsen, 2007 -- valid
	Cucochodaeus sparsus (LeConte, 1868) -- valid
	Cyclocephala Dejean, 1821 -- valid
	Cyclocephala aravaipensis Ratcliffe, 1992 -- valid
	Cyclocephala arenosa Howden and Endr�di, 1966 -- valid
	Cyclocephala barrerai Mart�nez, 1969 -- valid
	Cyclocephala borealis Arrow, 1911 -- valid
	Cyclocephala caelestis Delgado and Ratcliffe, 1990 -- valid
	Cyclocephala comata Bates, 1888 -- valid
	Cyclocephala deceptor (Casey, 1915) -- valid
	Cyclocephala dimidiata Burmeister, 1847 -- invalid
	Cyclocephala elegans Horn, 1871 -- invalid
	Cyclocephala forcipulata Howden and Endr�di, 1966 -- valid
	Cyclocephala freudei Endr�di, 1963 -- valid
	Cyclocephala grandis Burmeister, 1847 -- invalid
	Cyclocephala hirta LeConte, 1861 -- valid
	Cyclocephala knobelae (Brown, 1934) -- valid
	Cyclocephala landini Endr�di, 1964 -- invalid
	Cyclocephala longula LeConte, 1863 -- valid
	Cyclocephala lunulata Burmeister, 1847 -- valid
	Cyclocephala lurida Bland, 1863 -- valid
	Cyclocephala lurida coahuilae Bates, 1888 -- valid
	Cyclocephala lurida lurida Bland, 1863 -- valid
	Cyclocephala mafaffa Burmeister, 1847 -- valid
	Cyclocephala melanocephala (Fabricius, 1775) -- valid
	Cyclocephala mexica Mart�nez, 1969 -- invalid
	Cyclocephala miamiensis Howden and Endr�di, 1966 -- valid -- Miami roundhead scarab
	Cyclocephala nigricollis Burmeister, 1847 -- valid
	Cyclocephala nubeculosa Burmeister, 1847 -- invalid
	Cyclocephala ovulum Bates, 1888 -- invalid
	Cyclocephala parallela (Casey, 1915) -- valid
	Cyclocephala pasadenae (Casey, 1915) -- valid
	Cyclocephala pilosicollis Saylor, 1936 -- valid
	Cyclocephala puberula LeConte, 1863 -- valid
	Cyclocephala regularis Casey, 1915 -- valid
	Cyclocephala robusta LeConte, 1863 -- invalid
	Cyclocephala sanguinicollis Burmeister, 1847 -- valid
	Cyclocephala seditiosa LeConte, 1863 -- valid
	Cyclocephala sinaloae Howden and Endr�di, 1966 -- valid
	Cyclocephala sororia Bates, 1888 -- valid
	Cyclocephala sparsa Arrow, 1902 -- valid
	Cyclocephala testacea Burmeister, 1847 -- valid
	Cyclocephala variabilis Burmeister, 1847 -- valid
	Cyclocephala ventralis Erichson, 1847 -- invalid
	Cyclocephala villosa Burmeister, 1847 -- invalid
	Cyclocephala virkkii Howden and Endr�di, 1966 -- invalid
	Cyclocephala wandae Hardy, 1974 -- valid
	Cyclocephalini Laporte, 1840 -- valid
	Cyriopertha Reitter, 1903 -- invalid
	Dasydera LeConte, 1861 -- invalid
	Delipnia Casey, 1915 -- invalid
	Dellacasiellus Gordon and Skelley, 2007 -- valid
	Dellacasiellus carpinterius Gordon and Skelley, 2007 -- valid
	Dellacasiellus claudus (Fall, 1932) -- valid
	Dellacasiellus concavus (Say, 1823) -- valid
	Dellacasiellus fucosus (Schmidt, 1922) -- valid
	Dellacasiellus glamisensis Gordon and Skelley, 2007 -- valid
	Dellacasiellus griffini Gordon and Skelley, 2007 -- valid
	Dellacasiellus humboldti Gordon and Skelley, 2007 -- valid
	Dellacasiellus kirni (Cartwright, 1944) -- valid
	Dellacasiellus laevigatus (Haldeman, 1848) -- valid
	Dellacasiellus milleri Gordon and Skelley, 2007 -- valid
	Dellacasiellus pseudofucosus Gordon and Skelley, 2007 -- valid
	Dellacasiellus ruficlarus (Fall, 1932) -- valid
	Deltochilum Eschscholtz, 1822 -- valid
	Deltochilum gibbosum (Fabricius, 1775) -- valid
	Deltochilum gibbosum gibbosum (Fabricius, 1775) -- valid
	Deltochilum scabriusculum Bates, 1887 -- valid
	Deltochilum scabriusculum montanum Howden, 1966 -- valid
	Deltochilum scabriusculum scabriusculum Bates, 1887 -- valid
	Deltochilum tumidum Howden, 1966 -- valid
	Dialytellus Brown, 1929 -- valid
	Dialytellus dialytoides (Fall, 1907) -- valid
	Dialytellus tragicus (Schmidt, 1916) -- valid
	Dialytes Harold, 1869 -- valid
	Dialytes criddlei Brown, 1929 -- valid
	Dialytes striatulus (Say, 1825) -- valid
	Dialytes truncatus (Melsheimer, 1845) -- valid
	Dialytes ulkei Horn, 1875 -- valid
	Dialytodius Gordon and Skelley, 2007 -- valid
	Dialytodius carri (Brown, 1929) -- valid
	Dialytodius decipiens (Horn, 1887) -- valid
	Dialytodius olympicus Gordon and Skelley, 2007 -- valid
	Dialytodius rogersi (Hatch, 1971) -- valid
	Diapatalia Casey, 1915 -- invalid
	Diapterna Horn, 1887 -- valid
	Diapterna hamata (Say, 1824) -- valid
	Diapterna hyperborea (LeConte, 1850) -- valid
	Diapterna omissa (LeConte, 1850) -- valid
	Diapterna pinguella (Brown, 1929) -- valid
	Diapterna pinguis (Haldeman, 1848) -- valid
	Diastictus californicus Petrovitz, 1963 -- invalid
	Diazus LeConte, 1859 -- invalid
	Dichelonychini Burmeister, 1855 -- valid
	Dichelonyx Harris, 1827 -- valid
	Dichelonyx albicollis Burmeister, 1855 -- valid
	Dichelonyx arizonensis Saylor, 1933 -- invalid
	Dichelonyx arizonica Barrett, 1933 -- invalid
	Dichelonyx backii Kirby, 1837 -- valid
	Dichelonyx canadensis Horn, 1876 -- valid
	Dichelonyx clypeata Horn, 1876 -- valid
	Dichelonyx columbiana Hopping, 1931 -- invalid
	Dichelonyx crotchii Horn, 1876 -- invalid
	Dichelonyx crumbi Hatch, 1971 -- valid
	Dichelonyx decolorata Fall, 1901 -- valid
	Dichelonyx deserta Hopping, 1931 -- invalid
	Dichelonyx diluta Fall, 1901 -- valid
	Dichelonyx elongatula (Sch�nherr, 1817) -- valid
	Dichelonyx fulgida LeConte, 1856 -- invalid
	Dichelonyx fuscula LeConte, 1856 -- valid
	Dichelonyx kirbyi Brown, 1946 -- valid
	Dichelonyx lateralis Fall, 1901 -- valid
	Dichelonyx longiclava Fall, 1901 -- invalid
	Dichelonyx mormona Fall, 1907 -- invalid
	Dichelonyx muscula Fall, 1901 -- valid
	Dichelonyx nana Fall, 1901 -- valid
	Dichelonyx oregona Van Dyke, 1918 -- invalid
	Dichelonyx pallens LeConte, 1856 -- valid
	Dichelonyx picea Horn, 1894 -- valid
	Dichelonyx pusilla LeConte, 1856 -- valid
	Dichelonyx robusta Fall, 1901 -- valid
	Dichelonyx subvittata LeConte, 1856 -- valid
	Dichelonyx sulcata LeConte, 1856 -- valid
	Dichelonyx testacea Kirby, 1837 -- invalid
	Dichelonyx testaceipennis Fall, 1907 -- valid
	Dichelonyx truncata LeConte, 1856 -- valid
	Dichelonyx vaga Fall, 1901 -- valid
	Dichelonyx valida LeConte, 1856 -- valid
	Dichelonyx vandykei Saylor, 1932 -- valid
	Dichelonyx vicina Fall, 1901 -- valid
	Dichelonyx virescens Kirby, 1837 -- invalid
	Dichotomius Hope, 1838 -- valid
	Dichotomius carolinus (Linnaeus, 1767) -- valid
	Dichotomius colonicus (Say, 1835) -- valid
	Dichromina Casey, 1915 -- invalid
	Dichromina ocularis Casey, 1915 -- invalid
	Didactyliini Pittino, 1985 -- valid
	Dimalia Mulsant and Rey, 1870 -- invalid
	Dinacoma Casey, 1889 -- valid
	Dinacoma caseyi Blaisdell, 1930 -- valid -- Casey's June beetle
	Dinacoma marginata (Casey, 1886) -- valid
	Diphydactylus Thomson, 1858 -- invalid
	Diphyllostoma Fall, 1901 -- valid
	Diphyllostoma fimbriatum (Fall, 1901) -- valid
	Diphyllostoma linsleyi Fall, 1932 -- valid
	Diphyllostoma nigricollis Fall, 1912 -- valid
	Diphyllostomatidae Holloway, 1972 -- valid
	Diplotaxini Kirby, 1837 -- valid
	Diplotaxis Kirby, 1837 -- valid
	Diplotaxis abnormis Fall, 1909 -- valid
	Diplotaxis academia Vaurie, 1960 -- valid
	Diplotaxis acerba Fall, 1909 -- invalid
	Diplotaxis acononica Cazier, 1940 -- invalid
	Diplotaxis aenea Blanchard, 1851 -- valid
	Diplotaxis aequalis Cazier, 1940 -- valid
	Diplotaxis ambigua Fall, 1909 -- valid
	Diplotaxis angularis LeConte, 1856 -- valid
	Diplotaxis anthracina Fall, 1909 -- valid
	Diplotaxis anxia (LeConte, 1856) -- valid
	Diplotaxis arctifrons Bates, 1888 -- valid
	Diplotaxis arcuata Fall, 1909 -- invalid
	Diplotaxis arizonica Schaeffer, 1907 -- valid
	Diplotaxis arta Cazier, 1940 -- invalid
	Diplotaxis atlantis Fall, 1909 -- valid
	Diplotaxis atratula LeConte, 1856 -- valid
	Diplotaxis aulacochela Cazier, 1940 -- valid
	Diplotaxis australis Vaurie, 1960 -- valid
	Diplotaxis bakeri Cazier, 1940 -- valid
	Diplotaxis barbarae Vaurie and Cazier, 1955 -- valid
	Diplotaxis basalis Fall, 1909 -- valid
	Diplotaxis belfragei Fall, 1909 -- valid
	Diplotaxis beyeri Schaeffer, 1907 -- valid
	Diplotaxis bidentata LeConte, 1856 -- valid
	Diplotaxis blanchardi Vaurie, 1956 -- valid
	Diplotaxis boops Bates, 1888 -- valid
	Diplotaxis bowditchi Fall, 1909 -- valid
	Diplotaxis brachyptera Vaurie, 1960 -- valid
	Diplotaxis brevicollis LeConte, 1856 -- valid
	Diplotaxis brevicornis Cazier, 1940 -- valid
	Diplotaxis brevidens LeConte, 1856 -- valid
	Diplotaxis brevisetosa Linell, 1896 -- valid
	Diplotaxis caelestis Delgado and Capistran, 1993 -- valid
	Diplotaxis californica Schaeffer, 1907 -- invalid
	Diplotaxis carbonaria Burmeister, 1855 -- invalid
	Diplotaxis carbonata LeConte, 1856 -- valid
	Diplotaxis cartwrighti Cazier, 1940 -- invalid
	Diplotaxis castanea Burmeister, 1855 -- invalid
	Diplotaxis catarinas Vaurie and Cazier, 1955 -- valid
	Diplotaxis cephalotes Fall, 1909 -- invalid
	Diplotaxis chiricahuae Fall, 1909 -- valid
	Diplotaxis circulans Vaurie, 1960 -- valid
	Diplotaxis coenonychoides Davidson and Davidson, 2006 -- valid
	Diplotaxis commixta Vaurie, 1960 -- valid
	Diplotaxis compacta Fall, 1909 -- invalid
	Diplotaxis completa Cazier, 1940 -- valid
	Diplotaxis conformis Fall, 1909 -- valid
	Diplotaxis confusa Fall, 1909 -- valid
	Diplotaxis connata Schaeffer, 1905 -- valid
	Diplotaxis consors LeConte, 1856 -- invalid
	Diplotaxis contracta Bates, 1888 -- valid
	Diplotaxis convexilabra Vaurie and Cazier, 1955 -- valid
	Diplotaxis corbula Vaurie, 1960 -- valid
	Diplotaxis coriacea Bates, 1888 -- valid
	Diplotaxis corpulenta Burmeister, 1855 -- invalid
	Diplotaxis corvina LeConte, 1856 -- valid
	Diplotaxis costanera Vaurie, 1958 -- valid
	Diplotaxis costulata Fall, 1909 -- invalid
	Diplotaxis cribatella Bates, 1889 -- valid
	Diplotaxis cribulosa LeConte, 1856 -- valid
	Diplotaxis cribulosa cribulosa LeConte, 1856 -- valid
	Diplotaxis cribulosa sinaloa Vaurie, 1958 -- valid
	Diplotaxis curvaticeps Fall, 1909 -- valid
	Diplotaxis dahli Cazier, 1940 -- valid
	Diplotaxis decima Vaurie and Cazier, 1955 -- valid
	Diplotaxis densicollis Fall, 1909 -- invalid
	Diplotaxis dentella Fall, 1909 -- valid
	Diplotaxis denticeps Bates, 1887 -- valid
	Diplotaxis deserta Fall, 1932 -- valid
	Diplotaxis dissona Vaurie, 1960 -- valid
	Diplotaxis dubia LeConte, 1856 -- valid
	Diplotaxis durango Vaurie, 1960 -- valid
	Diplotaxis ebenina Blanchard, 1851 -- invalid
	Diplotaxis elongata Vaurie, 1960 -- valid
	Diplotaxis ennea Vaurie and Cazier, 1955 -- invalid
	Diplotaxis errans Fall, 1909 -- valid
	Diplotaxis excavata LeConte, 1856 -- invalid
	Diplotaxis exstans Vaurie, 1960 -- valid
	Diplotaxis falli Saylor, 1935 -- invalid
	Diplotaxis fimbriata Fall, 1909 -- valid
	Diplotaxis fissilabris Fall, 1909 -- valid
	Diplotaxis fissilis Vaurie and Cazier, 1955 -- valid
	Diplotaxis flexa Vaurie, 1960 -- valid
	Diplotaxis fossipalpa Fall, 1909 -- valid
	Diplotaxis frondicola (Say, 1825) -- valid
	Diplotaxis frontalis LeConte, 1856 -- invalid
	Diplotaxis fulgida Cazier, 1940 -- invalid
	Diplotaxis fulva (LeConte, 1856) -- valid
	Diplotaxis fulva Robinson, 1946 -- invalid
	Diplotaxis fusca Moser, 1918 -- invalid
	Diplotaxis futilis Fall, 1932 -- invalid
	Diplotaxis georgiae Blanchard, 1851 -- invalid
	Diplotaxis glabrimargo Vaurie and Cazier, 1955 -- valid
	Diplotaxis harperi Blanchard, 1851 -- valid
	Diplotaxis haydenii LeConte, 1856 -- valid
	Diplotaxis hebes Bates, 1888 -- valid
	Diplotaxis hirsuta Vaurie, 1958 -- valid
	Diplotaxis hispida Fall, 1909 -- invalid
	Diplotaxis illustris Fall, 1909 -- valid
	Diplotaxis impar Vaurie and Cazier, 1955 -- valid
	Diplotaxis impressicollis Moser, 1918 -- invalid
	Diplotaxis impressifrons Cazier, 1940 -- invalid
	Diplotaxis incuria Fall, 1932 -- valid
	Diplotaxis indigena Vaurie and Cazier, 1955 -- valid
	Diplotaxis ingenua Fall, 1909 -- valid
	Diplotaxis innoxia LeConte, 1856 -- invalid
	Diplotaxis insignis LeConte, 1861 -- valid
	Diplotaxis knausii Schaeffer, 1907 -- valid
	Diplotaxis kuschel Vaurie, 1960 -- valid
	Diplotaxis labrata Moser, 1918 -- invalid
	Diplotaxis laeviscutata Moser, 1918 -- invalid
	Diplotaxis languida LeConte, 1878 -- valid
	Diplotaxis lengii Fall, 1909 -- valid
	Diplotaxis levicosta Fall, 1909 -- valid
	Diplotaxis levicoxa Fall, 1909 -- invalid
	Diplotaxis levicula Casey, 1885 -- invalid
	Diplotaxis liberta (Germar, 1824) -- valid
	Diplotaxis macronycha Fall, 1909 -- valid
	Diplotaxis magna Bates, 1888 -- valid
	Diplotaxis marginicollis Fall, 1909 -- valid
	Diplotaxis martinezi Delgado and Capistran, 1993 -- valid
	Diplotaxis mascula Vaurie, 1960 -- valid
	Diplotaxis maura Fall, 1909 -- valid
	Diplotaxis megapleura Vaurie, 1958 -- valid
	Diplotaxis mentalis Fall, 1909 -- valid
	Diplotaxis mexicana Moser, 1918 -- invalid
	Diplotaxis microps Cazier, 1940 -- invalid
	Diplotaxis mima Vaurie and Cazier, 1955 -- valid
	Diplotaxis mimosae Fall, 1909 -- valid
	Diplotaxis misella Fall, 1909 -- valid
	Diplotaxis missionaria Vaurie, 1960 -- valid
	Diplotaxis moerens LeConte, 1856 -- valid
	Diplotaxis moerens moerens LeConte, 1856 -- valid
	Diplotaxis moerens peninsularis Fall, 1909 -- valid
	Diplotaxis morula LeConte, 1856 -- invalid
	Diplotaxis muricata Schaeffer, 1907 -- valid
	Diplotaxis mus Fall, 1932 -- valid
	Diplotaxis nigra Cartwright, 1939 -- invalid
	Diplotaxis nigriventris Bates, 1887 -- valid
	Diplotaxis obregon Vaurie, 1958 -- valid
	Diplotaxis obscura LeConte, 1859 -- valid
	Diplotaxis ohausi Moser, 1921 -- valid
	Diplotaxis pacata LeConte, 1856 -- valid
	Diplotaxis pala Vaurie, 1960 -- valid
	Diplotaxis parallela Fall, 1909 -- valid
	Diplotaxis parkeri Cazier, 1940 -- invalid
	Diplotaxis parpolita Vaurie, 1960 -- valid
	Diplotaxis parvicollis Fall, 1909 -- valid
	Diplotaxis persisae Cazier, 1940 -- valid
	Diplotaxis pilipennis Moser, 1918 -- valid
	Diplotaxis pinalica Fall, 1932 -- valid
	Diplotaxis pinguescens Fall, 1909 -- invalid
	Diplotaxis planidens Fall, 1909 -- valid
	Diplotaxis polita Fall, 1909 -- valid
	Diplotaxis popino Casey, 1885 -- invalid
	Diplotaxis producta Fall, 1932 -- invalid
	Diplotaxis profunda Vaurie and Cazier, 1955 -- valid
	Diplotaxis prominens Fall, 1909 -- invalid
	Diplotaxis protensa Fall, 1932 -- invalid
	Diplotaxis puberula LeConte, 1863 -- valid
	Diplotaxis pubipes Schaeffer, 1907 -- valid
	Diplotaxis pumila Fall, 1909 -- valid
	Diplotaxis punctata LeConte, 1856 -- valid
	Diplotaxis punctatorugosa Blanchard, 1851 -- valid
	Diplotaxis puncticeps Moser, 1918 -- valid
	Diplotaxis punctipennis LeConte, 1856 -- valid
	Diplotaxis punctulata Horn, 1894 -- valid
	Diplotaxis pygidialis Vaurie, 1960 -- valid
	Diplotaxis recticantha Cazier, 1940 -- invalid
	Diplotaxis residua Fall, 1909 -- valid
	Diplotaxis rex Vaurie, 1958 -- valid
	Diplotaxis robertmarki Davidson and Davidson, 2006 -- valid
	Diplotaxis rockefelleri Vaurie and Cazier, 1955 -- valid
	Diplotaxis rosae Vaurie and Cazier, 1955 -- valid
	Diplotaxis rotunda Vaurie, 1960 -- valid
	Diplotaxis rudis (LeConte, 1859) -- valid
	Diplotaxis rufa Linell, 1895 -- valid
	Diplotaxis rufiola Fall, 1909 -- valid
	Diplotaxis rufocastanea Moser, 1918 -- invalid
	Diplotaxis rugosioides Schaeffer, 1907 -- invalid
	Diplotaxis saltensis Vaurie, 1960 -- valid
	Diplotaxis saylori Cazier, 1940 -- valid
	Diplotaxis schaefferi Fall, 1909 -- valid
	Diplotaxis semifoveata Fall, 1909 -- valid
	Diplotaxis sericeiventris Moser, 1918 -- invalid
	Diplotaxis sierrae Fall, 1909 -- valid
	Diplotaxis simplex Blanchard, 1851 -- valid
	Diplotaxis sinuata Fall, 1909 -- invalid
	Diplotaxis sinuaticeps Bates, 1888 -- invalid
	Diplotaxis sonora Vaurie, 1960 -- valid
	Diplotaxis sordida (Say, 1825) -- valid
	Diplotaxis sparsa Vaurie, 1960 -- valid
	Diplotaxis sparsesetosa Schaeffer, 1907 -- invalid
	Diplotaxis stabilis Cazier, 1940 -- invalid
	Diplotaxis statura Cazier, 1940 -- valid
	Diplotaxis subangulata LeConte, 1856 -- valid
	Diplotaxis subcostata Blanchard, 1851 -- valid
	Diplotaxis subrugata Moser, 1918 -- valid
	Diplotaxis sulcatula Fall, 1909 -- valid
	Diplotaxis superflua Vaurie, 1960 -- valid
	Diplotaxis tarsalis Schaeffer, 1907 -- valid
	Diplotaxis tenebrosa Fall, 1909 -- valid
	Diplotaxis tenuis LeConte, 1856 -- valid
	Diplotaxis testacea Burmeister, 1855 -- invalid
	Diplotaxis texana LeConte, 1856 -- valid
	Diplotaxis thoracica Fall, 1909 -- valid
	Diplotaxis transversicollis Moser, 1918 -- invalid
	Diplotaxis trapezifera Bates, 1887 -- valid
	Diplotaxis trementina Fall, 1909 -- valid
	Diplotaxis tristis Kirby, 1837 -- valid
	Diplotaxis truncatula LeConte, 1856 -- valid
	Diplotaxis ungula Cazier, 1940 -- invalid
	Diplotaxis ungulata Cazier, 1940 -- valid
	Diplotaxis urbana Vaurie, 1960 -- valid
	Diplotaxis vandykei Vaurie, 1958 -- valid
	Diplotaxis villosa Fall, 1909 -- invalid
	Diplotaxis villosipes Fall, 1932 -- invalid
	Diplotaxis volatica Cazier, 1940 -- invalid
	Dorcus MacLeay, 1819 -- valid
	Dorcus brevis (Say, 1825) -- valid
	Dorcus carnochani Angell, 1916 -- invalid
	Dorcus caucasicus Ganglbauer, 1886 -- invalid
	Dorcus costatus LeConte, 1866 -- invalid
	Dorcus nanus Casey, 1909 -- invalid
	Dorcus parallelus (Say, 1824) -- valid
	Drepanacanthoides Schmidt, 1913 -- valid
	Drepanacanthoides larreae (Horn, 1887) -- valid
	Drepanacanthoides walshii (Horn, 1870) -- valid
	Dynastes MacLeay, 1819 -- valid
	Dynastes corniger Sternberg, 1910 -- invalid
	Dynastes granti Horn, 1870 -- valid
	Dynastes hyllus Chevrolat, 1843 -- valid
	Dynastes tityus (Linnaeus, 1767) -- valid
	Dynastinae MacLeay, 1819 -- valid
	Dynastini MacLeay, 1819 -- valid
	Dyscinetus Harold, 1869 -- valid
	Dyscinetus bitumorosus Casey, 1915 -- invalid
	Dyscinetus borealis Casey, 1915 -- invalid
	Dyscinetus discedens Casey, 1915 -- invalid
	Dyscinetus ebeninus Casey, 1915 -- invalid
	Dyscinetus geminatus Jacquelin du Val, 1856 -- invalid
	Dyscinetus gilanus Casey, 1915 -- invalid
	Dyscinetus laevicollis Arrow, 1937 -- valid
	Dyscinetus laevissimus Casey, 1915 -- invalid
	Dyscinetus morator (Fabricius, 1798) -- valid
	Dyscinetus obsidianus Casey, 1915 -- invalid
	Dyscinetus parvus Casey, 1915 -- invalid
	Dyscinetus picipes (Burmeister, 1847) -- valid
	Dyscinetus puncticauda Casey, 1909 -- invalid
	Dyscinetus punctipes Bates, 1888 -- invalid
	Dyscinetus subquadratus Casey, 1915 -- invalid
	Echmatophorus Waterhouse, 1895 -- invalid
	Ectinoplectron Ohaus, 1915 -- valid
	Ectinoplectron oryctoides (Ohaus, 1905) -- valid
	Endosa LeConte, 1856 -- invalid
	Endrosa volvula LeConte, 1856 -- invalid
	Enema Hope, 1837 -- valid
	Enema endymion Chevrolat, 1843 -- valid
	Enema gibbicollis Sternberg, 1908 -- invalid
	Enema lupercus Burmeister, 1847 -- invalid
	Enema paniscus Burmeister, 1847 -- invalid
	Eoglaresis Semenov-Tian-Shanskii and Medvedev, 1932 -- invalid
	Epichalcoplethis Bates, 1904 -- invalid
	Epipertinax Kuwert, 1891 -- invalid
	Epiphanus Kaup, 1871 -- invalid
	Epiphileurus Kolbe, 1910 -- invalid
	Epiphileurus septentrionis Kolbe, 1910 -- invalid
	Epiphoroneus Arrow, 1907 -- invalid
	Epipleurothrix Zang, 1905 -- invalid
	Eremoglaresis Semenov-Tian-Shanskii and Medvedev, 1932 -- invalid
	Erirhipidia Casey, 1915 -- invalid
	Erirhipis Burmeister, 1842 -- invalid
	Erirhipis clarkii LeConte, 1854 -- invalid
	Erirhipis occidentalis Knaus, 1916 -- invalid
	Eucanthus Westwood, 1848 -- valid
	Eucanthus alutaceus Cartwright, 1944 -- valid
	Eucanthus greeni Robinson, 1948 -- valid
	Eucanthus impressus Howden, 1964 -- valid
	Eucanthus lazarus (Fabricius, 1775) -- valid
	Eucanthus mexicanus Howden, 1964 -- valid
	Eucanthus subtropicus Howden, 1955 -- valid
	Euchlora maculata Laporte, 1840 -- invalid
	Euchlora nigritula Laporte, 1840 -- invalid
	Euclidius Kirby, 1827 -- invalid
	Euetheola Bates, 1888 -- valid
	Euetheola hondurana Casey, 1915 -- invalid
	Euetheola humilis (Burmeister, 1847) -- valid
	Euetheola subglabra (Schaeffer, 1909) -- valid
	Eugastra LeConte, 1856 -- invalid
	Euoniticellus Janssens, 1953 -- valid
	Euoniticellus cubiensis (Laporte, 1840) -- valid
	Euoniticellus intermedius (Reiche, 1849) -- valid
	Euparia LePeletier and Serville, 1828 -- valid
	Euparia castanea LePeletier and Serville, 1828 -- valid
	Eupariini Schmidt, 1910 -- valid
	Euparixia Brown, 1927 -- valid
	Euparixia duncani Brown, 1927 -- valid
	Euparixia mexicana Gordon and McCleve, 2003 -- valid
	Euparixia moseri Woodruff and Cartwright, 1967 -- valid
	Euphorhipis Casey, 1915 -- invalid
	Euphoria Burmeister, 1842 -- valid
	Euphoria aestuosa Horn, 1880 -- invalid
	Euphoria anneae (Howden, 1955) -- valid
	Euphoria appalachia Casey, 1915 -- invalid
	Euphoria areata (Fabricius, 1775) -- valid
	Euphoria arizonica Schaeffer, 1907 -- invalid
	Euphoria basalis (Gory and Percheron, 1833) -- valid
	Euphoria biguttata (Gory and Percheron, 1833) -- valid
	Euphoria binoculata Casey, 1915 -- invalid
	Euphoria biplagiata Casey, 1915 -- invalid
	Euphoria bispinis (Bates, 1889) -- valid
	Euphoria canescens (Gory and Percheron, 1833) -- valid
	Euphoria casselberryi Robinson, 1937 -- valid
	Euphoria connivens Casey, 1915 -- invalid
	Euphoria crinicauda Casey, 1915 -- invalid
	Euphoria crinitula Casey, 1915 -- invalid
	Euphoria cuprascens Casey, 1915 -- invalid
	Euphoria devulsa Horn, 1880 -- valid
	Euphoria discicollis (Thomson, 1878) -- valid
	Euphoria fascifera (LeConte, 1861) -- valid
	Euphoria fascifera fascifera (LeConte, 1861) -- valid
	Euphoria fascifera trapezium Casey, 1915 -- valid
	Euphoria floridana Casey, 1915 -- invalid
	Euphoria fulgida (Fabricius, 1775) -- valid
	Euphoria fulgida fulgida (Fabricius, 1775) -- valid
	Euphoria fulgida fuscocyanea Casey, 1915 -- valid
	Euphoria fulgida holochloris Fall, 1905 -- valid
	Euphoria fulgida limbalis Fall, 1905 -- valid
	Euphoria geminata (Chevrolat, 1835) -- valid
	Euphoria herbacea (Olivier, 1789) -- valid
	Euphoria hirtipes Horn, 1880 -- valid
	Euphoria histrionella Schaeffer, 1907 -- invalid
	Euphoria histrionica Thomson, 1878 -- valid
	Euphoria inda (Linnaeus, 1758) -- valid
	Euphoria kansana Casey, 1915 -- invalid
	Euphoria kernii Haldeman, 1852 -- valid
	Euphoria leucopyge Bates, 1889 -- valid
	Euphoria limatula Janson, 1881 -- invalid
	Euphoria lineoligera (Blanchard, 1850) -- valid
	Euphoria monticola Bates, 1889 -- valid
	Euphoria nigripennis Klages, 1894 -- invalid
	Euphoria oxysternum Casey, 1915 -- invalid
	Euphoria pilipennis (Kraatz, 1883) -- valid
	Euphoria pulchella (Gory and Percheron, 1833) -- valid
	Euphoria quadricollis Bates, 1889 -- valid
	Euphoria retusa Casey, 1915 -- invalid
	Euphoria rufobrunnea Casey, 1915 -- invalid
	Euphoria scabiosa Casey, 1915 -- invalid
	Euphoria schottii (LeConte, 1854) -- valid
	Euphoria scolopacea Casey, 1915 -- invalid
	Euphoria sepulchralis (Fabricius, 1801) -- valid
	Euphoria sepulchralis leucographa (Gory and Percheron, 1833) -- valid
	Euphoria sepulchralis nitens Casey, 1915 -- valid
	Euphoria sepulchralis rufina (Gory and Percheron, 1833) -- valid
	Euphoria sepulchralis sepulchralis (Fabricius, 1801) -- valid
	Euphoria solidula Casey, 1915 -- invalid
	Euphoria sonorae Bates, 1889 -- invalid
	Euphoria submetallica Casey, 1915 -- invalid
	Euphoria testacea Casey, 1915 -- invalid
	Euphoria texana Schaufuss, 1863 -- invalid
	Euphoria verticalis Horn, 1880 -- valid
	Euphoria wichitana Casey, 1915 -- invalid
	Euphoria xanthomelas Thomson, 1878 -- invalid
	Euphoria yucateca Bates, 1889 -- invalid
	Euphoriaspis Casey, 1915 -- invalid
	Euphoriopsis Casey, 1915 -- invalid
	Eupleurus Mulsant, 1842 -- valid
	Eupleurus subterraneus (Linnaeus, 1758) -- valid
	Eupopillia Kolbe, 1894 -- invalid
	Euryaspis Blanchard, 1851 -- invalid
	Exomala Reitter, 1903 -- invalid
	Flaviellus Gordon and Skelley, 2007 -- valid
	Flaviellus aggregatus (Gordon, 1977) -- valid
	Flaviellus gordoni (Ratcliffe, 1988) -- valid
	Flaviellus perfimbriatus (Gordon, 1977) -- valid
	Flaviellus phalerioides (Horn, 1870) -- valid
	Flaviellus phalerius Gordon and Skelley, 2007 -- valid
	Flaviellus subtruncatus (LeConte, 1878) -- valid
	Fossocarus Howden, 1961 -- valid
	Fossocarus creoleorum Howden, 1961 -- valid
	Gananota Ohaus, 1915 -- invalid
	Genuchinus Westwood, 1874 -- valid
	Genuchinus angustus Casey, 1915 -- invalid
	Genuchinus grandis Van Dyke, 1952 -- invalid
	Genuchinus ineptus (Horn, 1885) -- valid
	Geocanthon Pereira and Mart�nez, 1956 -- invalid
	Geomyphilus Gordon and Skelley, 2007 -- valid
	Geomyphilus alabama (Skelley and Gordon, 1995) -- valid
	Geomyphilus essigi (Saylor, 1935) -- valid
	Geomyphilus geronimo Gordon and Skelley, 2007 -- valid
	Geomyphilus insolitus (Brown, 1928) -- valid
	Geomyphilus kiowensis (Gordon and Salsbury, 1999) -- valid
	Geomyphilus leptotarsis (Brown, 1928) -- valid
	Geomyphilus pinolus Gordon and Skelley, 2007 -- valid
	Geomyphilus rubiginosus (Horn, 1870) -- valid
	Geomyphilus russeus (Brown, 1928) -- valid
	Geomyphilus thomomysi (Brown, 1928) -- valid
	Geomyphilus ungulatus (Fall, 1901) -- valid
	Geomyphilus viceversus (Paulsen, 2006) -- valid
	Geopsammodius Gordon and Pittino, 1992 -- valid
	Geopsammodius fuscus Skelley, 2006 -- valid
	Geopsammodius hydropicus (Horn, 1887) -- valid
	Geopsammodius morrisi Skelley, 2006 -- valid
	Geopsammodius ohoopee Skelley, 2006 -- valid
	Geopsammodius relictillus (Deyrup and Woodruff, 1991) -- valid
	Geopsammodius rileyi Skelley, 2006 -- valid
	Geopsammodius subpedalis Skelley, 2006 -- valid
	Geopsammodius unsidensis Skelley, 2006 -- valid
	Geopsammodius withlacoochee Skelley, 2006 -- valid
	Geotrupes Latreille, 1797 -- valid
	Geotrupes (Anoplotrupes) Jekel, 1865 -- valid
	Geotrupes (Cnemotrupes) Jekel, 1865 -- valid
	Geotrupes (Geohowdenius) Zunino, 1984 -- valid
	Geotrupes (Geotrupes) Latreille, 1797 -- valid
	Geotrupes (Megatrupes) Zunino, 1984 -- valid
	Geotrupes aeneus Felsche, 1909 -- invalid
	Geotrupes balyi Jekel, 1865 -- valid
	Geotrupes blackburnii (Fabricius, 1781) -- valid
	Geotrupes blackburnii blackburnii (Fabricius, 1781) -- valid
	Geotrupes blackburnii excrementi Say, 1823 -- valid
	Geotrupes cavicollis Bates, 1887 -- valid
	Geotrupes chalybaeus LeConte, 1878 -- invalid
	Geotrupes chevrolati Jekel, 1865 -- invalid
	Geotrupes conicollis Jekel, 1865 -- invalid
	Geotrupes egeriei Germar, 1824 -- valid
	Geotrupes gilnickii Jekel, 1865 -- invalid
	Geotrupes haldemani Jekel, 1865 -- invalid
	Geotrupes hornii Blanchard, 1888 -- valid
	Geotrupes jekelii Horn, 1868 -- invalid
	Geotrupes lecontei Jekel, 1865 -- invalid
	Geotrupes melsheimeri Jekel, 1865 -- invalid
	Geotrupes mixtus Horn, 1868 -- invalid
	Geotrupes opacus Haldeman, 1853 -- valid
	Geotrupes ovalipennis Jekel, 1865 -- invalid
	Geotrupes semiopacus Jekel, 1865 -- valid
	Geotrupes similis Jekel, 1865 -- invalid
	Geotrupes splendidulus (Fabricius, 1775) -- valid
	Geotrupes splendidulus miarophagus Say, 1823 -- valid
	Geotrupes splendidulus splendidulus (Fabricius, 1775) -- valid
	Geotrupes starki Jekel, 1865 -- invalid
	Geotrupes stercorarius (Linnaeus, 1758) -- valid
	Geotrupes ulkei Blanchard, 1888 -- valid
	Geotrupidae Latreille, 1802 -- valid
	Geotrupinae Latreille, 1802 -- valid
	Germarostes Paulian, 1982 -- valid
	Germarostes (Germarostes) Paulian, 1982 -- valid
	Germarostes aphodioides (Illiger, 1800) -- valid
	Germarostes globosus (Say, 1835) -- valid
	Gillaspytes Howden, 1980 -- valid
	Gillaspytes janzeni Howden, 1980 -- valid
	Glaphyridae MacLeay, 1819 -- valid
	Glaphyrinae MacLeay, 1819 -- valid
	Glaresidae Kolbe, 1905 -- valid
	Glaresis Erichson, 1848 -- valid
	Glaresis arenata Gordon, 1974 -- valid -- Kelso Dune glaresis scarab
	Glaresis canadensis Brown, 1928 -- valid
	Glaresis cartwrighti Gordon, 1970 -- valid
	Glaresis clypeata Van Dyke, 1928 -- valid
	Glaresis confusa Brown, 1928 -- valid
	Glaresis dakotensis Gordon, 1970 -- valid
	Glaresis ecostata Fall, 1907 -- valid
	Glaresis gordoni Warner, 1995 -- valid
	Glaresis howdeni Gordon, 1970 -- valid
	Glaresis inducta Horn, 1885 -- valid
	Glaresis knausi Brown, 1928 -- invalid
	Glaresis medialis Gordon, 1969 -- valid
	Glaresis mendica Horn, 1885 -- valid
	Glaresis phoenicis Fall, 1907 -- valid
	Glaresis texana Gordon, 1970 -- valid
	Glaresis zvirgzdinsi Warner, 1995 -- valid
	Gnomon Zang, 1905 -- invalid
	Gnorimella Casey, 1915 -- valid
	Gnorimella maculosa (Knoch, 1801) -- valid
	Gnorimus dissimilis Gory and Percheron, 1833 -- invalid
	Godschama Reitter, 1903 -- invalid
	Golofa Hope, 1837 -- valid
	Golofa (Golofa) Hope, 1837 -- valid
	Golofa imperialis Thomson, 1858 -- valid
	Golofa xiximeca Mor�n, 1995 -- valid
	Gonocyphus Lansberge, 1885 -- invalid
	Goraqua P�ringuey, 1907 -- invalid
	Goraqua smithsana P�ringuey, 1907 -- invalid
	Graphalia Casey, 1915 -- invalid
	Gronocarus Schaeffer, 1927 -- valid
	Gronocarus autumnalis Schaeffer, 1927 -- valid
	Gronocarus inornatus Skelley, 2005 -- valid
	Gronocarus multispinosus Howden, 1961 -- invalid -- Spiny Florida sandhill scarab
	Gymnetina Kirby, 1827 -- valid
	Gymnetina Casey, 1915 -- valid
	Gymnetina cretacea (LeConte, 1866) -- valid
	Gymnetini Kirby, 1827 -- valid
	Gymnetis MacLeay, 1819 -- valid
	Gymnetis aequalis Casey, 1915 -- invalid
	Gymnetis atrata Gory and Percheron, 1833 -- invalid
	Gymnetis caseyi Antoine, 2001 -- valid
	Gymnetis cuneata Casey, 1915 -- invalid
	Gymnetis laetula Casey, 1915 -- invalid
	Gymnetis lobiculata Casey, 1915 -- invalid
	Gymnetis mexicana Gory and Percheron, 1833 -- invalid
	Gymnetis nigrorubra Gory and Percheron, 1833 -- invalid
	Gymnetis palliata Gory and Percheron, 1833 -- invalid
	Gymnetis punctata Blanchard, 1850 -- invalid
	Gymnetis simulans Casey, 1915 -- invalid
	Gymnetis sobrina Gory and Percheron, 1833 -- invalid
	Gymnetis uniformis Blanchard, 1850 -- invalid
	Gymnetoides Mart�nez, 1949 -- invalid
	Gymnetosoma Mart�nez, 1949 -- invalid
	Gymnodus Kirby, 1827 -- invalid
	Gymnopyge Linell, 1896 -- valid
	Gymnopyge coquilletti Linell, 1896 -- valid
	Gymnopyge hirsuta Cazier, 1939 -- valid
	Gymnopyge hopliaeformis Linell, 1896 -- valid
	Gymnopyge pygmaea Linell, 1896 -- valid
	Gymnoropterus Gestro, 1899 -- invalid
	Gynnis LeConte, 1856 -- invalid
	Halffterinetis Mor�n and Nogueira, 2007 -- valid
	Halffterinetis gonzaloi Mor�n and Nogueira, 2007 -- valid
	Halffterinetis violetae Mor�n and Nogueira, 2007 -- valid
	Halotosia Casey, 1915 -- invalid
	Haplophoria Casey, 1915 -- invalid
	Haroldiataenius Chalumeau, 1981 -- valid
	Haroldiataenius buvexus Stebnicka and Skelley, 2009 -- valid
	Haroldiataenius convexus (Robinson, 1940) -- valid
	Haroldiataenius griffini (Cartwright, 1974) -- valid
	Haroldiataenius hintoni (Saylor, 1933) -- valid
	Haroldiataenius limbatus (Bates, 1887) -- valid
	Haroldiataenius lucanus (Horn, 1871) -- valid
	Haroldiataenius mariarum (Bates, 1887) -- valid
	Haroldiataenius saramari (Cartwright, 1939) -- valid
	Haroldiataenius semipilosus (Van Dyke, 1928) -- valid
	Haroldiellus Gordon and Skelley, 2007 -- valid
	Haroldiellus sallei (Harold, 1863) -- valid
	Heliscus Zang, 1906 -- valid
	Heliscus tropicus (Percheron, 1835) -- valid
	Hemicyclorhinus Seabra, 1909 -- invalid
	Hemiphileurus Kolbe, 1910 -- valid
	Hemiphileurus illatus (LeConte, 1854) -- valid
	Hemispilota Casey, 1915 -- invalid
	Heteronychus tumulosus Burmeister, 1847 -- invalid
	Hexalus Mulsant and Rey, 1870 -- invalid
	Hexaphyllus Mulsant, 1839 -- invalid
	Hologymnetis Mart�nez, 1949 -- valid
	Hologymnetis argenteola (Bates, 1889) -- valid
	Hologymnetis cinerea (Gory and Percheron, 1833) -- valid
	Homochromina Casey, 1915 -- invalid
	Homocopris Burmeister, 1842 -- invalid
	Homoiosternus Ohaus, 1901 -- valid
	Homoiosternus beckeri Ohaus, 1901 -- valid
	Homovalgus Kolbe, 1897 -- invalid
	Hoplia Illiger, 1803 -- valid
	Hoplia (Hoplia) Illiger, 1803 -- valid
	Hoplia barbata Blatchley, 1910 -- invalid
	Hoplia callipyge LeConte, 1856 -- valid
	Hoplia cazieri Boyer, 1940 -- invalid
	Hoplia cinereonebulosa Nonfried, 1895 -- valid
	Hoplia convexula LeConte, 1856 -- invalid
	Hoplia debilis LeConte, 1856 -- invalid
	Hoplia deserticola Boyer, 1940 -- invalid
	Hoplia dispar LeConte, 1880 -- valid
	Hoplia equina LeConte, 1880 -- valid
	Hoplia ferrisi Boyer, 1940 -- invalid
	Hoplia floridana Fisher, 1918 -- valid
	Hoplia gilleti Hardy, 1977 -- valid
	Hoplia helvola Melsheimer, 1845 -- invalid
	Hoplia hirta LeConte, 1880 -- valid
	Hoplia humboltensis Boyer, 1940 -- invalid
	Hoplia irrorata LeConte, 1856 -- invalid
	Hoplia laticollis LeConte, 1856 -- valid
	Hoplia lecontei Dalla Torre, 1913 -- invalid
	Hoplia meridionalis Boyer, 1940 -- invalid
	Hoplia modesta Haldeman, 1843 -- valid
	Hoplia monticola Melsheimer, 1845 -- invalid
	Hoplia mucorea (Germar, 1824) -- valid
	Hoplia mutata Harold, 1869 -- invalid
	Hoplia oregona LeConte, 1856 -- invalid
	Hoplia primaria Burmeister, 1844 -- invalid
	Hoplia pubicollis LeConte, 1856 -- invalid
	Hoplia sackenii LeConte, 1880 -- valid
	Hoplia singularis Burmeister, 1844 -- invalid
	Hoplia trifasciata Say, 1825 -- valid
	Hoplia tristis Melsheimer, 1845 -- invalid
	Hoplia trivialis Harold, 1869 -- valid
	Hoplia utahensis Boyer, 1940 -- invalid
	Hopliatus Rafinesque, 1815 -- invalid
	Hopliini Latreille, 1829 -- valid
	Hornietus Stebnicka, 2000 -- valid
	Hornietus ventralis (Horn, 1887) -- valid
	Hyboma LePeletier and Serville, 1828 -- invalid
	Hybosoridae Erichson, 1847 -- valid
	Hybosorinae Erichson, 1847 -- valid
	Hybosorus MacLeay, 1819 -- valid
	Hybosorus carolinus LeConte, 1848 -- invalid
	Hybosorus illigeri Reiche, 1853 -- invalid
	Hybosorus nitidus Lansberge, 1882 -- invalid
	Hybosorus nossibianus Fairmaire, 1895 -- invalid
	Hybosorus palearcticus Endr�di, 1957 -- invalid
	Hybosorus pinguis Westwood, 1845 -- invalid
	Hybosorus roei Westwood, 1845 -- valid
	Hypothyce Howden, 1968 -- valid
	Hypothyce burnei Skelley, 2005 -- valid
	Hypothyce mixta Howden, 1968 -- valid
	Hypothyce osburni (Cartwright, 1967) -- valid
	Hypotrichia LeConte, 1861 -- valid
	Hypotrichia spissipes LeConte, 1861 -- valid
	Inca LePeletier and Serville, 1828 -- valid
	Inca clathrata sommeri Westwood, 1845 -- valid
	Incaini Burmeister, 1842 -- invalid
	Incini Burmeister, 1842 -- valid
	Irrasinus Gordon and Skelley, 2007 -- valid
	Irrasinus dumasi Gordon and Skelley, 2007 -- valid
	Irrasinus stupidus (Horn, 1870) -- valid
	Isocoryna Casey, 1915 -- invalid
	Isonychus Mannerheim, 1829 -- valid
	Isonychus arizonensis Howden, 1959 -- valid
	Isorhipina Casey, 1915 -- invalid
	Junkia Dalla Torre, 1913 -- invalid
	Kolbeus Boucomont, 1911 -- invalid
	Labarrus Mulsant and Rey, 1870 -- valid
	Labarrus lividus (Olivier, 1789) -- valid
	Labarrus pseudolividus Balthasar, 1941 -- valid
	Lachnosterna Hope, 1837 -- invalid
	Lachnosterna affinis LeConte, 1856 -- invalid
	Lachnosterna alpina Linell, 1896 -- invalid
	Lachnosterna anodentata Bates, 1888 -- invalid
	Lachnosterna arcuata Smith, 1888 -- invalid
	Lachnosterna biimpresa Smith, 1889 -- invalid
	Lachnosterna boops Horn, 1887 -- invalid
	Lachnosterna burmeisteri LeConte, 1856 -- invalid
	Lachnosterna carolina Fall, 1912 -- invalid
	Lachnosterna cephalica LeConte, 1856 -- invalid
	Lachnosterna cerasina LeConte, 1856 -- invalid
	Lachnosterna ciliata LeConte, 1856 -- invalid
	Lachnosterna consimilis LeConte, 1850 -- invalid
	Lachnosterna cribricollis Bates, 1888 -- invalid
	Lachnosterna dampfi Arrow, 1933 -- invalid
	Lachnosterna decidua LeConte, 1856 -- invalid
	Lachnosterna definita Smith, 1889 -- invalid
	Lachnosterna dubia Smith, 1888 -- invalid
	Lachnosterna exorata Horn, 1887 -- invalid
	Lachnosterna frontalis LeConte, 1856 -- invalid
	Lachnosterna generosa Horn, 1887 -- invalid
	Lachnosterna georgiana Schaeffer, 1909 -- invalid
	Lachnosterna glabripennis LeConte, 1856 -- invalid
	Lachnosterna grandior Linell, 1896 -- invalid
	Lachnosterna grandis Smith, 1888 -- invalid
	Lachnosterna guatemalica Moser, 1918 -- invalid
	Lachnosterna heterodoxa Horn, 1887 -- invalid
	Lachnosterna hirticeps LeConte, 1856 -- invalid
	Lachnosterna inana LeConte, 1856 -- invalid
	Lachnosterna innominata Smith, 1889 -- invalid
	Lachnosterna insperata Smith, 1889 -- invalid
	Lachnosterna integra LeConte, 1856 -- invalid
	Lachnosterna lenta Fall, 1908 -- invalid
	Lachnosterna limula Horn, 1887 -- invalid
	Lachnosterna longiclava Moser, 1918 -- invalid
	Lachnosterna longipilosa Bates, 1888 -- invalid
	Lachnosterna lugubris LeConte, 1856 -- invalid
	Lachnosterna lutescens LeConte, 1856 -- invalid
	Lachnosterna minor Linell, 1896 -- invalid
	Lachnosterna nitidula LeConte, 1863 -- invalid
	Lachnosterna nova Smith, 1889 -- invalid
	Lachnosterna obesa LeConte, 1856 -- invalid
	Lachnosterna parilis Bates, 1889 -- invalid
	Lachnosterna parva Linell, 1896 -- invalid
	Lachnosterna politula Horn, 1887 -- invalid
	Lachnosterna pygidialis Schaeffer, 1906 -- invalid
	Lachnosterna quadrata Smith, 1889 -- invalid
	Lachnosterna robusta LeConte, 1856 -- invalid
	Lachnosterna rufiola LeConte, 1856 -- invalid
	Lachnosterna rugosioides Linell, 1896 -- invalid
	Lachnosterna semicribrata LeConte, 1856 -- invalid
	Lachnosterna serricornis LeConte, 1856 -- invalid
	Lachnosterna setipennis Bates, 1888 -- invalid
	Lachnosterna sororia LeConte, 1856 -- invalid
	Lagopelas Burmeister, 1876 -- invalid
	Lasioperix Zang, 1905 -- invalid
	Lasiopus LeConte, 1856 -- invalid
	Latemnis Thomson, 1880 -- invalid
	Lechorodius Gordon and Skelley, 2007 -- valid
	Lechorodius lutulentus (Haldeman, 1843) -- valid
	Lechorodius terminalis (Say, 1823) -- valid
	Leiopsammodius Rakovic, 1981 -- valid
	Leiopsammodius acei Harpootlian, Gordon and Woodruff, 2000 -- valid
	Leiopsammodius deyrupi Harpootlian, Gordon and Woodruff, 2000 -- valid
	Leiopsammodius malkini (Cartwright, 1946) -- valid
	Leiopsammodius ocmulgeei Harpootlian, Gordon and Woodruff, 2000 -- valid
	Leptaegialia Brown, 1931 -- invalid
	Leptohoplia Saylor, 1935 -- valid
	Leptohoplia testaceipennis Saylor, 1935 -- valid
	Leptohopliina Potts, 1974 -- valid
	Liatongus Reitter, 1893 -- valid
	Liatongus californicus (Horn, 1882) -- valid
	Lichnanthe Burmeister, 1844 -- valid
	Lichnanthe albipilosa Carlson, 1980 -- valid -- White sand bear scarab
	Lichnanthe apina Carlson, 1980 -- valid
	Lichnanthe brachyselis Carlson, 1980 -- valid
	Lichnanthe canina Horn, 1867 -- invalid
	Lichnanthe cooperi (Horn, 1867) -- valid
	Lichnanthe edwardsi Horn, 1870 -- invalid
	Lichnanthe lupina LeConte, 1856 -- valid
	Lichnanthe rathvoni (LeConte, 1863) -- valid
	Lichnanthe ursina (LeConte, 1861) -- valid -- Bumblebee scarab
	Lichnanthe vulpina (Hentz, 1827) -- valid
	Ligniperda Fabricius, 1790 -- invalid
	Ligyrellus Casey, 1915 -- invalid
	Ligyrodes Casey, 1915 -- invalid
	Ligyrodes aztecus Casey, 1915 -- invalid
	Ligyrodes clypealis Casey, 1915 -- invalid
	Ligyrodes dawsoni Casey, 1924 -- invalid
	Ligyrodes parviceps Casey, 1915 -- invalid
	Ligyrodes propinquus Casey, 1915 -- invalid
	Ligyrodes quadripennis Casey, 1915 -- invalid
	Ligyrodes vernicicollis Casey, 1915 -- invalid
	Ligyrus Burmeister, 1847 -- invalid
	Ligyrus arizonensis Casey, 1915 -- invalid
	Ligyrus bicorniculatus Casey, 1915 -- invalid
	Ligyrus blatchleyi Cartwright, 1944 -- invalid
	Ligyrus brevipes Casey, 1915 -- invalid
	Ligyrus breviusculus Casey, 1915 -- invalid
	Ligyrus bryanti Rivers, 1891 -- invalid
	Ligyrus californicus Casey, 1909 -- invalid
	Ligyrus curtipennis Casey, 1915 -- invalid
	Ligyrus effetus Casey, 1915 -- invalid
	Ligyrus farctus Casey, 1915 -- invalid
	Ligyrus lacustris Casey, 1915 -- invalid
	Ligyrus laetulus Casey, 1915 -- invalid
	Ligyrus laevicauda Casey, 1915 -- invalid
	Ligyrus laticauda Casey, 1915 -- invalid
	Ligyrus laticollis Casey, 1915 -- invalid
	Ligyrus longulus Casey, 1915 -- invalid
	Ligyrus lucublandus Casey, 1915 -- invalid
	Ligyrus parallelus Casey, 1915 -- invalid
	Ligyrus puncticauda Casey, 1915 -- invalid
	Ligyrus pygidialis Bates, 1888 -- invalid
	Ligyrus remotus Casey, 1915 -- invalid
	Ligyrus rubidus Casey, 1915 -- invalid
	Ligyrus rugiceps LeConte, 1856 -- invalid
	Ligyrus scitulus Casey, 1915 -- invalid
	Ligyrus spissipes Casey, 1909 -- invalid
	Ligyrus texanus Casey, 1915 -- invalid
	Ligyrus virginicus Casey, 1915 -- invalid
	Limaroides Balthasar, 1932 -- invalid
	Liogenys pauperata Burmeister, 1855 -- invalid
	Liothorax Motschulsky, 1859 -- valid
	Liothorax alternatus (Horn, 1870) -- valid
	Liothorax consociatus (Horn, 1887) -- valid
	Liothorax innexus (Say, 1835) -- valid
	Liothorax subaeneus (LeConte, 1857) -- valid
	Lissomelas Bates, 1889 -- valid
	Lissomelas flohri Bates, 1889 -- valid
	Listrochelus debilis Arrow, 1933 -- invalid
	Listrochelus durangoensis Bates, 1888 -- invalid
	Listrochelus gracilis Horn, 1885 -- invalid
	Listrochelus langeri Chapin, 1934 -- invalid
	Listrochelus longiclavus Fall, 1922 -- invalid
	Listrochelus obtusus LeConte, 1856 -- invalid
	Listrochelus puberulus LeConte, 1863 -- invalid
	Listrochelus testaceipennis Moser, 1921 -- invalid
	Listrochelus texanus LeConte, 1856 -- invalid
	Litocopris Waterhouse, 1891 -- invalid
	Lonchophorus Germar, 1824 -- invalid
	Lophocephalus Kuwert, 1896 -- invalid
	Lucanidae Latreille, 1804 -- valid -- stag beetles
	Lucaninae Latreille, 1804 -- valid
	Lucanini Latreille, 1804 -- valid
	Lucanus Scopoli, 1763 -- valid
	Lucanus capreolus (Linnaeus, 1764) -- valid -- stag beetle
	Lucanus carlengi Angell, 1916 -- invalid
	Lucanus dama Fabricius, 1775 -- invalid
	Lucanus elaphus Fabricius, 1775 -- valid
	Lucanus lentus Laporte, 1840 -- invalid
	Lucanus mazama (LeConte, 1861) -- valid
	Lucanus muticus Thunberg, 1806 -- invalid
	Lucanus oblongus de Charpentier, 1825 -- invalid
	Lucanus placidus Say, 1825 -- valid
	Lucanus quercus Weber, 1801 -- invalid
	Lucanus trigonus Thunberg, 1806 -- invalid
	Lucilius Kuwert, 1891 -- invalid
	Luxolinus Gordon and Skelley, 2007 -- valid
	Luxolinus luxatus (Horn, 1887) -- valid
	Lycophontes Bruch, 1910 -- invalid
	Macranoxia Crotch, 1873 -- invalid
	Macrodactylini Kirby, 1837 -- valid
	Macrodactylus Dejean, 1821 -- valid
	Macrodactylus angustatus (Palisot de Beauvois, 1805) -- valid
	Macrodactylus angustatus LeConte, 1856 -- invalid
	Macrodactylus barbatus Fitch, 1863 -- invalid
	Macrodactylus fulvescens Bates, 1887 -- valid
	Macrodactylus impressus Bates, 1887 -- valid
	Macrodactylus infuscatus Bates, 1887 -- valid
	Macrodactylus lineaticollis Bates, 1887 -- valid
	Macrodactylus mexicanus Burmeister, 1855 -- valid
	Macrodactylus murinus Bates, 1887 -- valid
	Macrodactylus nigripes Bates, 1887 -- valid
	Macrodactylus nigritarsis Bates, 1887 -- invalid
	Macrodactylus polyphagus Burmeister, 1855 -- invalid
	Macrodactylus rhomboderus Bates, 1887 -- invalid
	Macrodactylus rufescens Bates, 1887 -- valid
	Macrodactylus setulosus LeConte, 1856 -- invalid
	Macrodactylus silaonus Bates, 1887 -- valid
	Macrodactylus subspinosus (Fabricius, 1775) -- valid
	Macrodactylus uniformis Horn, 1876 -- valid
	Macrodactylus variipes Bates, 1887 -- valid
	Macrodactylus vicinus Bates, 1887 -- invalid
	Macrodactylus virens Bates, 1887 -- valid
	Macrolobus Zang, 1905 -- invalid
	Maculaphodius Gordon and Skelley, 2007 -- valid
	Maculaphodius conspersus (Horn, 1887) -- valid
	Maladera Mulsant and Rey, 1871 -- valid
	Maladera (Cephaloserica) Brenske, 1900 -- valid
	Maladera castanea (Arrow, 1913) -- valid
	Maladera castanea castanea (Arrow, 1913) -- valid
	Malagoniella Mart�nez, 1961 -- valid
	Malagoniella astyanax yucateca (Harold, 1863) -- valid
	Manlius Kuwert, 1891 -- invalid
	Martinezia Chalumeau, 1983 -- invalid
	Martineziana Chalumeau and Ozdikmen, 2006 -- valid
	Martineziana dutertrei (Chalumeau, 1983) -- valid
	Martineziana vandykei (Hinton, 1936) -- valid
	Martineziella Chalumeau, 1987 -- invalid
	Megalisus Mulsant and Rey, 1870 -- invalid
	Megalosoma Burmeister, 1847 -- invalid
	Megalotrox De Borre, 1886 -- invalid
	Megapertha Reitter, 1903 -- invalid
	Megasoma Kirby, 1825 -- valid
	Megasoma cedrosum Hardy, 1972 -- valid
	Megasoma lecontei Hardy, 1972 -- valid
	Megasoma lenczyi Cartwright, 1976 -- valid
	Megasoma occidentalis Bol�var y Pieltain, Jim�nez-As�a and Mart�nez, 1963 -- valid
	Megasoma pachecoi Cartwright, 1963 -- valid
	Megasoma punctulatum Cartwright, 1952 -- valid
	Megasoma sleeperi Hardy, 1972 -- valid
	Megasoma thersites LeConte, 1861 -- valid
	Megasoma vogti Cartwright, 1963 -- valid
	Megasominus Casey, 1915 -- invalid
	Meghyboma Kolbe, 1893 -- invalid
	Melanocanthon Halffter, 1958 -- valid
	Melanocanthon bispinatus (Robinson, 1941) -- valid
	Melanocanthon granulifer (Schmidt, 1920) -- valid
	Melanocanthon nigricornis (Say, 1823) -- valid
	Melanocanthon punctaticollis (Schaeffer, 1915) -- valid
	Melanotrupes Blanchard, 1888 -- invalid
	Melinopterus Mulsant, 1842 -- valid
	Melinopterus consentaneus (LeConte, 1850) -- valid
	Melinopterus femoralis (Say, 1823) -- valid
	Melinopterus prodromus (Brahm, 1790) -- valid
	Melolontha annulata Germar, 1824 -- invalid
	Melolontha aphodiina Billberg, 1820 -- invalid
	Melolontha atrata Fabricius, 1801 -- invalid
	Melolontha elongata Fabricius, 1793 -- invalid
	Melolontha elongata Herbst, 1790 -- invalid
	Melolontha fervens Gyllenhal, 1817 -- invalid
	Melolontha georgicana Gyllenhal, 1817 -- invalid
	Melolontha hexagona Germar, 1824 -- invalid
	Melolontha leucogramma Blanchard, 1851 -- invalid
	Melolontha linearis Gyllenhal, 1817 -- invalid
	Melolontha lutea Olivier, 1789 -- invalid
	Melolontha minima Drapiez, 1819 -- invalid
	Melolontha moerens Fabricius, 1801 -- invalid
	Melolontha moesta Say, 1825 -- invalid
	Melolontha pilosicollis Knoch, 1801 -- invalid
	Melolontha porcina Hentz, 1830 -- invalid
	Melolontha quadrimaculata Drapiez, 1820 -- invalid
	Melolontha quercina Knoch, 1801 -- invalid
	Melolontha rufescens Latreille, 1804 -- invalid
	Melolontha semirufa Gyllenhal, 1817 -- invalid
	Melolontha unifasciata Say, 1825 -- invalid
	Melolonthinae Leach, 1819 -- valid
	Melolonthini Leach, 1819 -- valid
	Mendidius Harold, 1868 -- valid
	Mendidius aculeatus (Robinson, 1940) -- valid
	Merogyrus Gordon and Skelley, 2007 -- valid
	Merogyrus rotundiceps (Fall, 1907) -- valid
	Metapopillia Kolbe, 1894 -- invalid
	Micraegialia Brown, 1931 -- valid
	Micraegialia pusilla (Horn, 1887) -- valid
	Microcopris Balthasar, 1958 -- invalid
	Microphylla quedenfeldtii Kraatz, 1894 -- invalid
	Mitrorhinus Kaup, 1871 -- invalid
	Monapus Erichson, 1848 -- invalid
	Mononidia Casey, 1915 -- invalid
	Morosophus Kuwert, 1896 -- invalid
	Mycotrupes LeConte, 1866 -- valid
	Mycotrupes cartwrighti Olson and Hubbell, 1954 -- valid
	Mycotrupes gaigei Olson and Hubbell, 1954 -- valid
	Mycotrupes lethroides (Westwood, 1837) -- valid
	Mycotrupes pedester Howden, 1954 -- valid -- Scrub Island burrowing scarab
	Mycotrupes retusus (LeConte, 1866) -- valid
	Myrmeceicon Mann, 1914 -- invalid
	Nefoncerus Saylor, 1938 -- valid
	Nefoncerus convergens (Horn, 1894) -- valid
	Neleides Kaup, 1869 -- invalid
	Neleurops Kuwert, 1891 -- invalid
	Neleus acarinatus Kuwert, 1898 -- invalid
	Neleus aequatoris Kuwert, 1898 -- invalid
	Neleus altidens Kuwert, 1898 -- invalid
	Neleus approximatidentatus Kuwert, 1898 -- invalid
	Neleus arcuatotaeniatus Kuwert, 1898 -- invalid
	Neleus argentinus Kuwert, 1898 -- invalid
	Neleus arrogans Kuwert, 1898 -- invalid
	Neleus boliviensis Kuwert, 1898 -- invalid
	Neleus carinaefrons Kuwert, 1898 -- invalid
	Neleus chilensis Kuwert, 1891 -- invalid
	Neleus cognettii Pangella, 1905 -- invalid
	Neleus difficilis Kuwert, 1891 -- invalid
	Neleus difformis Kuwert, 1898 -- invalid
	Neleus dilatidentatus Kuwert, 1898 -- invalid
	Neleus dilatipunctatus Kuwert, 1898 -- invalid
	Neleus dilatus Kuwert, 1898 -- invalid
	Neleus disjunctus Kuwert, 1898 -- invalid
	Neleus dislocandus Kuwert, 1898 -- invalid
	Neleus dispar Kuwert, 1898 -- invalid
	Neleus dispositus Kuwert, 1898 -- invalid
	Neleus distinctus Kuwert, 1898 -- invalid
	Neleus distinguendus Kuwert, 1891 -- invalid
	Neleus durangi Kuwert, 1898 -- invalid
	Neleus festae Rosmini, 1902 -- invalid
	Neleus guatemalae Kuwert, 1891 -- invalid
	Neleus intermissus Kuwert, 1898 -- invalid
	Neleus laborator Kaup, 1869 -- invalid
	Neleus labroexcisus Kuwert, 1898 -- invalid
	Neleus laeviclypeatus Kuwert, 1891 -- invalid
	Neleus nicaraguae Kuwert, 1891 -- invalid
	Neleus obtusecornutus Kuwert, 1898 -- invalid
	Neleus sanio Kuwert, 1891 -- invalid
	Neleus scelus Kuwert, 1891 -- invalid
	Neleus scepticus Kuwert, 1891 -- invalid
	Neleus scurra Kuwert, 1891 -- invalid
	Neleus scurroides Kuwert, 1898 -- invalid
	Neleus scutellosulcatus Kuwert, 1898 -- invalid
	Neleus subcarinaefrons Kuwert, 1898 -- invalid
	Neleus subcarinatus Kuwert, 1898 -- invalid
	Neleus sulcicornis Kuwert, 1898 -- invalid
	Neleus taeniolatus Kuwert, 1898 -- invalid
	Neleus transvalensis Kuwert, 1891 -- invalid
	Neleus vagans Kuwert, 1891 -- invalid
	Neoathyreus Howden and Mart�nez, 1963 -- valid
	Neoathyreus fissicornis (Harold, 1880) -- valid
	Neoathyreus mixtus Howden, 1964 -- valid
	Neochodaeus Nikolayev, 1995 -- valid
	Neochodaeus frontalis (LeConte, 1863) -- valid
	Neochodaeus praesidii (Bates, 1887) -- valid
	Neochodaeus repandus (Fall, 1909) -- valid
	Neochodaeus striatus (LeConte, 1854) -- valid
	Neopsammodius Rakovic, 1986 -- valid
	Neopsammodius blandus (Fall, 1932) -- valid
	Neopsammodius interruptus (Say, 1835) -- valid
	Neopsammodius mimeticus (Fall, 1932) -- valid
	Neopsammodius quinqueplicatus (Horn, 1871) -- valid
	Neopsammodius saltilloensis (Cartwright, 1955) -- valid
	Neopsammodius werneri (Cartwright, 1955) -- valid
	Neotrichonotulus Dellacasa, Gordon and Dellacasa, 2004 -- valid
	Neotrichonotulus inurbanus (Gordon and Howden, 1973) -- valid
	Nialaphodius Kolbe, 1908 -- valid
	Nialaphodius nigritus (Fabricius, 1801) -- valid
	Nicagini LeConte, 1861 -- valid
	Nicagus LeConte, 1861 -- valid
	Nicagus obscurus (LeConte, 1847) -- valid
	Nicagus occultus Paulsen and Smith, 2005 -- valid
	Ninoides Kuwert, 1891 -- invalid
	Nipponoserica Nomura, 1973 -- valid
	Nipponoserica peregrina (Chapin, 1938) -- valid
	Ochodaeidae Mulsant and Rey, 1871 -- valid
	Ochodaeinae Mulsant and Rey, 1871 -- valid
	Ochodaeini Mulsant and Rey, 1871 -- valid
	Ochodaeus complex LeConte, 1868 -- invalid
	Ochodaeus kansanus Fall, 1909 -- invalid
	Ochodaeus mandibularis Linell, 1896 -- invalid
	Ochodaeus nimius Fall, 1907 -- invalid
	Ochodaeus opacus LeConte, 1868 -- invalid
	Ochrosidia Casey, 1915 -- invalid
	Ochrosidia abrupta Casey, 1915 -- invalid
	Ochrosidia ambiens Casey, 1915 -- invalid
	Ochrosidia arizonica Casey, 1915 -- invalid
	Ochrosidia californica Arrow, 1937 -- invalid
	Ochrosidia facilis Casey, 1915 -- invalid
	Ochrosidia marcida Casey, 1915 -- invalid
	Ochrosidia melina Casey, 1915 -- invalid
	Ochrosidia modulata Casey, 1915 -- invalid
	Ochrosidia obesula Casey, 1915 -- invalid
	Ochrosidia oblita Casey, 1915 -- invalid
	Ochrosidia oblongula Casey, 1915 -- invalid
	Ochrosidia ovulata Casey, 1915 -- invalid
	Ochrosidia pagana Casey, 1915 -- invalid
	Ochrosidia phasma Casey, 1915 -- invalid
	Ochrosidia prona Casey, 1915 -- invalid
	Ochrosidia protenta Casey, 1915 -- invalid
	Ochrosidia pusilla Casey, 1915 -- invalid
	Ochrosidia reflexa Casey, 1915 -- invalid
	Ochrosidia rufifrons Casey, 1915 -- invalid
	Ochrosidia rugulifrons Casey, 1915 -- invalid
	Ochrosidia rustica Casey, 1915 -- invalid
	Ochrosidia subvittata Brown, 1930 -- invalid
	Ochrosidia tenuicutis Casey, 1915 -- invalid
	Ochrosidia validiceps Casey, 1915 -- invalid
	Odonteus Samouelle, 1819 -- valid
	Odonteus alabamensis Wallis, 1929 -- valid
	Odonteus cornigerus (Melsheimer, 1845) -- valid
	Odonteus darlingtoni Wallis, 1928 -- valid
	Odonteus falli Wallis, 1928 -- valid
	Odonteus filicornis (Say, 1823) -- valid
	Odonteus floridensis Wallis, 1928 -- valid
	Odonteus liebecki Wallis, 1928 -- valid
	Odonteus obesus LeConte, 1859 -- valid
	Odonteus simi Wallis, 1928 -- valid
	Odonteus thoracicornis Wallis, 1928 -- valid
	Odontognathus Laporte, 1840 -- invalid
	Odontopsammodius Gordon and Pittino, 1992 -- valid
	Odontopsammodius armaticeps (Fall, 1932) -- valid
	Odontopsammodius cruentus (Harold, 1867) -- valid
	Odontotaenius Kuwert, 1896 -- valid
	Odontotaenius disjunctus (Illiger, 1800) -- valid -- bessbug
	Odontotaenius floridanus Schuster, 1994 -- valid
	Odontotaenius striatopunctatus (Percheron, 1835) -- valid
	Oeneus Kuwert, 1896 -- invalid
	Ohkubous Sawada, 1938 -- invalid
	Oliganomala Casey, 1915 -- invalid
	Omaloplia trogiformis Uhler, 1855 -- invalid
	Omorgus Erichson, 1847 -- valid
	Omorgus asper LeConte, 1854 -- valid
	Omorgus carinatus (Loomis, 1922) -- valid
	Omorgus fuliginosus (Robinson, 1941) -- valid
	Omorgus howelli (Howden and Vaurie, 1957) -- valid -- Caracara commensal scarab
	Omorgus inflatus (Loomis, 1922) -- valid
	Omorgus integer LeConte, 1854 -- invalid
	Omorgus monachus (Herbst, 1790) -- valid
	Omorgus morsus LeConte, 1854 -- invalid
	Omorgus nodosus (Robinson, 1940) -- valid
	Omorgus punctatus (Germar, 1824) -- valid
	Omorgus pustulatus LeConte, 1854 -- invalid
	Omorgus rubricans (Robinson, 1946) -- valid
	Omorgus scabrosus (Palisot de Beauvois, 1818) -- valid
	Omorgus scutellaris (Say, 1824) -- valid
	Omorgus suberosus (Fabricius, 1775) -- valid
	Omorgus suturalis LeConte, 1854 -- invalid
	Omorgus tesselatus LeConte, 1854 -- valid
	Omorgus texanus LeConte, 1854 -- valid
	Omorgus tomentosus (Robinson, 1941) -- valid
	Omorgus tytus (Robinson, 1941) -- valid
	Omorgus umbonatus LeConte, 1854 -- valid
	Oncerini LeConte, 1861 -- valid
	Oncerus LeConte, 1856 -- valid
	Oncerus floralis LeConte, 1856 -- valid
	Oniticellini Kolbe, 1905 -- valid
	Oniticellus LePeletier and Serville, 1828 -- valid
	Oniticellus rhinocerulus Bates, 1889 -- valid
	Onitini Laporte, 1840 -- valid
	Onitis Fabricius, 1798 -- valid
	Onitis alexis Klug, 1835 -- valid
	Ontherus Erichson, 1847 -- valid
	Ontherus (Caelontherus) G�nier, 1996 -- valid
	Ontherus mexicanus Harold, 1868 -- valid
	Onthophagini Burmeister, 1846 -- valid
	Onthophagus Latreille, 1802 -- valid
	Onthophagus (Digitonthophagus) Balthasar, 1959 -- valid
	Onthophagus (Onthophagus) Latreille, 1802 -- valid
	Onthophagus aciculatulus Blatchley, 1928 -- valid
	Onthophagus alluvius Howden and Cartwright, 1963 -- valid
	Onthophagus alternans d'Orbigny, 1898 -- invalid
	Onthophagus alternatus d'Orbigny, 1898 -- invalid
	Onthophagus alutaceus Blatchley, 1916 -- invalid
	Onthophagus antivagans Howden and G�nier, 2004 -- valid
	Onthophagus arizonensis Schaeffer, 1909 -- invalid
	Onthophagus arnetti Howden and Cartwright, 1963 -- valid
	Onthophagus athenae Goidanich, 1925 -- invalid
	Onthophagus batesi Howden and Cartwright, 1963 -- valid
	Onthophagus bos Villa and Villa, 1833 -- invalid
	Onthophagus bovillus Mulsant, 1842 -- invalid
	Onthophagus brevifrons Horn, 1881 -- valid
	Onthophagus browni Howden and Cartwright, 1963 -- valid
	Onthophagus canadensis Horn, 1875 -- invalid
	Onthophagus canelasensis Howden and G�nier, 2004 -- valid
	Onthophagus capreolus Mulsant, 1842 -- invalid
	Onthophagus carteri Blackburn, 1904 -- invalid
	Onthophagus cartwrighti Howden, 1973 -- valid
	Onthophagus castaneus Melsheimer, 1845 -- invalid
	Onthophagus castanonotus Seabra, 1907 -- invalid
	Onthophagus cavernicollis Howden and Cartwright, 1963 -- valid
	Onthophagus cervicornis Kirby, 1825 -- invalid
	Onthophagus chevrolati Harold, 1869 -- valid
	Onthophagus chevrolati chevrolati Harold, 1869 -- valid
	Onthophagus chevrolati longecarinatus Zunino and Halffter, 1988 -- valid
	Onthophagus circumcinctus Escalera, 1914 -- invalid
	Onthophagus coahuilae Zunino and Halffter, 1988 -- valid
	Onthophagus cochisus Brown, 1927 -- valid
	Onthophagus concinnus Laporte, 1840 -- valid
	Onthophagus concinnus Horn, 1875 -- invalid
	Onthophagus coproides Horn, 1881 -- valid
	Onthophagus cribricollis Horn, 1881 -- valid
	Onthophagus cuboidalis Bates, 1887 -- invalid
	Onthophagus cuevensis Howden, 1973 -- valid
	Onthophagus curvicornis Latreille, 1812 -- invalid
	Onthophagus cyanellus Bates, 1887 -- valid
	Onthophagus cynomysi Brown, 1927 -- valid
	Onthophagus depressus Harold, 1871 -- valid
	Onthophagus falcipes Harold, 1871 -- invalid
	Onthophagus femineus Mulsant, 1842 -- invalid
	Onthophagus fuscipennis Mulsant, 1842 -- invalid
	Onthophagus fuscus Boucomont, 1932 -- valid
	Onthophagus fuscus fuscus Boucomont, 1932 -- valid
	Onthophagus gazella (Fabricius, 1787) -- valid
	Onthophagus glareanus Gistel, 1857 -- invalid
	Onthophagus hecate (Panzer, 1794) -- valid
	Onthophagus hecate blatchleyi Brown, 1929 -- valid
	Onthophagus hecate hecate (Panzer, 1794) -- valid
	Onthophagus hidalgus Zunino and Halffter, 1988 -- valid
	Onthophagus hoepfneri Harold, 1869 -- valid
	Onthophagus incensus Say, 1835 -- valid
	Onthophagus intermedius Reiche, 1840 -- invalid
	Onthophagus knausi Brown, 1927 -- valid
	Onthophagus knulli Howden and Cartwright, 1963 -- valid
	Onthophagus landolti Harold, 1880 -- valid
	Onthophagus landolti landolti Harold, 1880 -- valid
	Onthophagus landolti texanus Schaeffer, 1914 -- valid
	Onthophagus lecontei Harold, 1871 -- valid
	Onthophagus mcclevei Howden and G�nier, 2004 -- valid
	Onthophagus medorensis Brown, 1929 -- valid
	Onthophagus mendax Mulsant, 1842 -- invalid
	Onthophagus mextexus Howden and Cartwright, 1970 -- valid
	Onthophagus monticulus Howden and Cartwright, 1963 -- invalid
	Onthophagus morio Brull�, 1832 -- invalid
	Onthophagus nigrovirescens Mulsant, 1842 -- invalid
	Onthophagus nuchicornis (Linnaeus, 1758) -- valid
	Onthophagus oklahomensis Brown, 1927 -- valid
	Onthophagus orpheus (Panzer, 1794) -- valid
	Onthophagus orpheus canadensis (Fabricius, 1801) -- valid
	Onthophagus orpheus orpheus (Panzer, 1794) -- valid
	Onthophagus orpheus pseudorpheus Howden and Cartwright, 1963 -- valid
	Onthophagus pennsylvanicus Harold, 1871 -- valid
	Onthophagus polyphemi Hubbard, 1894 -- valid -- Onthophagus tortoise commensal scarab
	Onthophagus polyphemi polyphemi Hubbard, 1894 -- valid
	Onthophagus polyphemi sparsisetosus Howden and Cartwright, 1963 -- valid
	Onthophagus protensus Melsheimer, 1845 -- invalid
	Onthophagus protensus Horn, 1875 -- invalid
	Onthophagus pseudofuscus Zunino and Halffter, 1988 -- valid
	Onthophagus reichardti Medvedev, 1930 -- invalid
	Onthophagus rhinoceros Melsheimer, 1845 -- invalid
	Onthophagus rubripennis Goidanich, 1924 -- invalid
	Onthophagus saharae Bedel, 1911 -- invalid
	Onthophagus sayi Laporte, 1840 -- invalid
	Onthophagus scabricollis Kirby, 1837 -- invalid
	Onthophagus scabricollis Horn, 1875 -- invalid
	Onthophagus schaefferi Howden and Cartwright, 1963 -- valid
	Onthophagus striatulus (Palisot de Beauvois, 1809) -- valid
	Onthophagus striatulus floridanus Blatchley, 1928 -- valid
	Onthophagus striatulus striatulus (Palisot de Beauvois, 1809) -- valid
	Onthophagus subaeneus Horn, 1875 -- invalid
	Onthophagus subaeneus (Palisot de Beauvois, 1811) -- valid
	Onthophagus subopacus Robinson, 1940 -- valid
	Onthophagus subtropicus Howden and Cartwright, 1963 -- valid
	Onthophagus taurus (Schreber, 1759) -- valid
	Onthophagus tuberculifrons Harold, 1871 -- valid
	Onthophagus velutinus Horn, 1875 -- valid
	Onthophagus viridicatus Say, 1835 -- invalid
	Onthophagus viridicollis Harold, 1869 -- invalid
	Onthurgus Gistel, 1857 -- invalid
	Onychotrupes Jekel, 1865 -- invalid
	Opiocanthon Paulian, 1947 -- invalid
	Orizabus Fairmaire, 1878 -- valid
	Orizabus batesi Prell, 1914 -- valid
	Orizabus brevicollis Prell, 1914 -- valid
	Orizabus clunalis (LeConte, 1856) -- valid
	Orizabus cultripes Fairmaire, 1878 -- invalid
	Orizabus fontinalis Casey, 1915 -- invalid
	Orizabus isodonoides Fairmaire, 1878 -- valid
	Orizabus ligyrodes Horn, 1885 -- invalid
	Orizabus marginatus Fairmaire, 1878 -- invalid
	Orizabus parvitarsis Casey, 1915 -- invalid
	Orizabus ponderosus Casey, 1915 -- invalid
	Orizabus pyriformis (LeConte, 1847) -- valid
	Orizabus ratcliffei Delgado, 2008 -- valid
	Orizabus rubricollis Prell, 1914 -- valid
	Orizabus sallei Fairmaire, 1878 -- invalid
	Orizabus snowi Horn, 1885 -- invalid
	Orizabus verticalis Fall, 1905 -- invalid
	Orsonyx LeConte, 1856 -- invalid
	Oryctini Mulsant, 1842 -- valid
	Oscarinus Gordon and Skelley, 2007 -- valid
	Oscarinus abusus (Fall, 1907) -- valid
	Oscarinus bottimeri (Cartwright, 1957) -- valid
	Oscarinus brimleyi (Cartwright, 1939) -- valid
	Oscarinus crassuloides (Fall, 1907) -- valid
	Oscarinus crassulus (Horn, 1870) -- valid
	Oscarinus floridanus (Robinson, 1947) -- valid
	Oscarinus lodingi (Cartwright, 1957) -- valid
	Oscarinus matiganae (Paulsen, 2006) -- valid
	Oscarinus odocoilis (Robinson, 1939) -- valid
	Oscarinus pseudabusus (Cartwright, 1957) -- valid
	Oscarinus rusicola (Melsheimer, 1845) -- valid
	Oscarinus silvanicus (Cartwright, 1972) -- valid
	Oscarinus stuessyi Gordon and Skelley, 2007 -- valid
	Oscarinus texensis (Cartwright, 1972) -- valid
	Oscarinus welderi Gordon and Skelley, 2007 -- valid
	Oscarinus windsori (Cartwright, 1939) -- valid
	Osmoderma LePeletier and Serville, 1828 -- valid
	Osmoderma beauvoisii Bainbridge, 1841 -- invalid
	Osmoderma caviceps Casey, 1915 -- invalid
	Osmoderma delicatula Casey, 1915 -- invalid
	Osmoderma eremicola (Knoch, 1801) -- valid
	Osmoderma gracilipes Casey, 1915 -- invalid
	Osmoderma lacustrina Casey, 1915 -- invalid
	Osmoderma montana Wickham, 1920 -- invalid
	Osmoderma scabra (Palisot de Beauvois, 1805) -- valid
	Osmoderma subplanata Casey, 1915 -- valid
	Osmodermatini Schenkling, 1922 -- valid
	Otophorus Mulsant, 1842 -- valid
	Otophorus haemorrhoidalis (Linnaeus, 1758) -- valid
	Oxygrylius Casey, 1915 -- valid
	Oxygrylius peninsularis Casey, 1915 -- valid
	Oxygrylius pimalis Casey, 1915 -- invalid
	Oxygrylius ruginasus (LeConte, 1856) -- valid
	Oxyomus Dejean, 1833 -- valid
	Oxyomus opacifrons Horn, 1871 -- invalid
	Oxyomus silvestris (Scopoli, 1763) -- valid
	Pachydemini Burmeister, 1855 -- valid
	Pachyplectrinae Ocampo, 2006 -- valid
	Pachyplectrus LeConte, 1874 -- valid
	Pachyplectrus laevis LeConte, 1874 -- valid
	Pachystethus Blanchard, 1851 -- invalid
	Pachystethus floridana Robinson, 1941 -- invalid
	Paedhyboma Kolbe, 1893 -- invalid
	Palaeocopris Pierce, 1946 -- invalid
	Palechus Casey, 1915 -- invalid
	Parabyrsopolis Ohaus, 1915 -- valid
	Parabyrsopolis chihuahuae (Bates, 1888) -- valid
	Parabyrsopolis villadealendensis Mor�n, 1976 -- invalid
	Parachrysina Bates, 1888 -- valid
	Parachrysina borealis Jameson, 1991 -- valid
	Paraclinidia Mart�nez, 1965 -- invalid
	Paracopris Balthasar, 1939 -- invalid
	Paracotalpa Ohaus, 1915 -- valid
	Paracotalpa deserta Saylor, 1940 -- valid
	Paracotalpa granicollis (Haldeman, 1852) -- valid
	Paracotalpa piceola Saylor, 1940 -- invalid
	Paracotalpa puncticollis (LeConte, 1863) -- valid
	Paracotalpa ursina (Horn, 1867) -- valid
	Paramaladera Nikolajev, 1979 -- invalid
	Paranomala Casey, 1915 -- invalid
	Parapertinax Kuwert, 1896 -- invalid
	Parapsammodius Gordon and Pittino, 1992 -- valid
	Parapsammodius bidens (Horn, 1871) -- valid
	Parapsammodius puncticollis (LeConte, 1858) -- valid
	Parareoda Casey, 1915 -- invalid
	Parareoda rufobrunnea Casey, 1915 -- invalid
	Parastasia Westwood, 1842 -- valid
	Parastasia brevipes (LeConte, 1856) -- valid
	Parataenius Balthasar, 1961 -- valid
	Parataenius granuliceps Petrovitz, 1971 -- invalid
	Parataenius simulator (Harold, 1868) -- valid
	Parathyce Hardy, 1974 -- invalid
	Pardalosus Gordon and Skelley, 2007 -- valid
	Pardalosus neodistinctus (Brown, 1928) -- valid
	Pardalosus pardalis (LeConte, 1857) -- valid
	Pardalosus pseudopardalis Gordon and Skelley, 2007 -- valid
	Pardalosus pumilio (Schmidt, 1907) -- valid
	Pardalosus sayi (Gordon, 2006) -- valid
	Pardalosus serval (Say, 1835) -- valid
	Pardalosus slevini (Van Dyke, 1928) -- valid
	Parisorhipis Casey, 1915 -- invalid
	Parochodaeus Nikolayev, 1995 -- valid
	Parochodaeus biarmatus (LeConte, 1863) -- valid
	Parochodaeus californicus (Horn, 1895) -- valid
	Parochodaeus duplex (LeConte, 1868) -- valid
	Parochodaeus howdeni (Carlson, 1975) -- valid
	Parochodaeus inarmatus (Schaeffer, 1906) -- valid
	Parochodaeus pectoralis (LeConte, 1868) -- valid
	Parochodaeus peninsularis (Horn, 1895) -- valid
	Parochodaeus ritcheri (Carlson, 1975) -- valid
	Passalidae Leach, 1815 -- valid -- bess beetles
	Passalinae Leach, 1815 -- valid
	Passalini Leach, 1815 -- valid
	Passalus Fabricius, 1792 -- valid
	Passalus bos Kuwert, 1891 -- invalid
	Passalus cornutus Fabricius, 1801 -- invalid
	Passalus distinctus Weber, 1801 -- invalid
	Passalus nodus Apetz, 1837 -- invalid
	Passalus punctatissimus Eschscholtz, 1829 -- invalid
	Passalus punctatostriatus Percheron, 1835 -- valid
	Passalus punctiger LePeletier and Serville, 1825 -- valid
	Passalus riograndensis Luederwaldt, 1931 -- valid
	Passalus striolatus Eschscholtz, 1829 -- invalid
	Passalus thoracicus Smith, 1852 -- invalid
	Pelidnota MacLeay, 1819 -- valid
	Pelidnota alutacea Bates, 1888 -- invalid
	Pelidnota brevicollis Casey, 1915 -- invalid
	Pelidnota brevis Casey, 1915 -- invalid
	Pelidnota cuprascens Casey, 1915 -- invalid
	Pelidnota debiliceps Casey, 1915 -- invalid
	Pelidnota howdeni Hardy, 1975 -- invalid
	Pelidnota hudsonica Casey, 1915 -- invalid
	Pelidnota laeta Sturm, 1843 -- invalid
	Pelidnota lucae LeConte, 1863 -- valid
	Pelidnota lugubris LeConte, 1874 -- valid
	Pelidnota oblonga Casey, 1915 -- invalid
	Pelidnota obscurella Casey, 1915 -- invalid
	Pelidnota ornatissima Sturm, 1843 -- invalid
	Pelidnota pallidipes Casey, 1915 -- invalid
	Pelidnota permicans Casey, 1915 -- invalid
	Pelidnota perplexa Hardy, 1975 -- valid
	Pelidnota planipennis Ohaus, 1918 -- invalid
	Pelidnota ponderella Casey, 1915 -- invalid
	Pelidnota punctata (Linnaeus, 1758) -- valid
	Pelidnota refulgens Casey, 1915 -- invalid
	Pelidnota strenua Casey, 1915 -- invalid
	Pelidnota strigosa Laporte, 1840 -- valid
	Pelidnota tarsalis Casey, 1915 -- invalid
	Pelidnota texensis Casey, 1915 -- invalid
	Pelidnota virescens Burmeister, 1844 -- valid
	Pelidnotidia Casey, 1915 -- invalid
	Pelidnotopsis Ohaus, 1915 -- invalid
	Peltotrupes Blanchard, 1888 -- valid
	Peltotrupes dubius Howden, 1955 -- invalid
	Peltotrupes profundus (Howden, 1952) -- valid
	Peltotrupes youngi Howden, 1955 -- valid -- Ocala burrowing scarab
	Pentodontini Mulsant, 1842 -- valid
	Periphileurus Kolbe, 1910 -- invalid
	Peripopillia Kolbe, 1894 -- invalid
	Pertinacides Kuwert, 1891 -- invalid
	Pertinax Kaup, 1869 -- invalid
	Petrejus Kaup, 1869 -- invalid
	Phaeaphodius Reitter, 1892 -- valid
	Phaeaphodius Schmidt, 1922 -- invalid
	Phaeaphodius rectus (Motschulsky, 1866) -- valid
	Phalangochaeta Mart�nez, 1952 -- invalid
	Phalangochaeta grandis Petrovitz, 1973 -- invalid
	Phanaeini Hope, 1838 -- valid
	Phanaeus MacLeay, 1819 -- valid
	Phanaeus (Phanaeus) MacLeay, 1819 -- valid
	Phanaeus adonis Harold, 1863 -- valid
	Phanaeus amethystinus Harold, 1863 -- valid
	Phanaeus amethystinus amethystinus Harold, 1863 -- valid
	Phanaeus amithaon Harold, 1875 -- valid
	Phanaeus borealis D'Olsoufieff, 1924 -- invalid
	Phanaeus cyanellus Robinson, 1938 -- invalid
	Phanaeus difformis LeConte, 1847 -- valid
	Phanaeus flohri Nevinson, 1892 -- valid
	Phanaeus floridanus D'Olsoufieff, 1924 -- invalid
	Phanaeus furcosus Felsche, 1901 -- invalid
	Phanaeus furiosus Bates, 1887 -- valid
	Phanaeus genieri Arnaud, 2001 -- valid
	Phanaeus goldanichi Balthasar, 1939 -- invalid
	Phanaeus igneus MacLeay, 1819 -- valid
	Phanaeus laevipennis Sturm, 1843 -- invalid
	Phanaeus magnificens Robinson, 1948 -- invalid
	Phanaeus martinezi Halffter, 1955 -- invalid
	Phanaeus niger D'Olsoufieff, 1924 -- invalid
	Phanaeus palliatus Sturm, 1843 -- valid
	Phanaeus quadridens (Say, 1835) -- valid
	Phanaeus rubervirens Robinson, 1948 -- invalid
	Phanaeus torrens LeConte, 1847 -- invalid
	Phanaeus triangularis (Say, 1823) -- valid
	Phanaeus triangularis texensis Edmonds, 1994 -- valid
	Phanaeus triangularis triangularis (Say, 1823) -- valid
	Phanaeus vindex MacLeay, 1819 -- valid
	Phanaeus violaceus Laporte, 1840 -- invalid
	Phanaeus yecoraensis Edmonds, 2004 -- valid
	Phaulothorax Zang, 1905 -- invalid
	Phileurini Burmeister, 1847 -- valid
	Phileurus Latreille, 1807 -- valid
	Phileurus capra Bates, 1888 -- invalid
	Phileurus carolinae Casey, 1915 -- invalid
	Phileurus castaneus Haldeman, 1843 -- invalid
	Phileurus floridanus Casey, 1915 -- invalid
	Phileurus recurvatus Casey, 1915 -- invalid
	Phileurus sulcifer Casey, 1915 -- invalid
	Phileurus texensis Casey, 1915 -- invalid
	Phileurus truncatus (Palisot de Beauvois, 1806) -- valid
	Phileurus vitulus LeConte, 1863 -- invalid
	Philochloenia Dejean, 1833 -- invalid
	Phobetus LeConte, 1856 -- valid
	Phobetus centralis Casey, 1909 -- invalid
	Phobetus chearyi Hardy, 1973 -- valid
	Phobetus ciliatus Barrett, 1935 -- valid
	Phobetus comatus LeConte, 1856 -- valid
	Phobetus desertus Blom and Clark, 1984 -- valid
	Phobetus humeralis Cazier, 1937 -- valid
	Phobetus leachi Barrett, 1935 -- invalid
	Phobetus mojavus Barrett, 1933 -- valid
	Phobetus palpalis Saylor, 1936 -- valid
	Phobetus panamintensis Hardy, 1978 -- valid
	Phobetus robinsoni Saylor, 1939 -- invalid
	Phobetus saylori Cazier, 1937 -- valid
	Phobetus sleeperi Hardy, 1978 -- valid
	Phobetus sloopi Barrett, 1933 -- invalid
	Phobetus testaceus LeConte, 1861 -- valid
	Phoronaeosomus Kuwert, 1891 -- invalid
	Phyllopertha Stephens, 1830 -- invalid
	Phyllopertha tanbaensis Nijima and Kinoshita, 1923 -- invalid
	Phyllopertha xanthogastra Harold, 1881 -- invalid
	Phyllophaga Harris, 1827 -- valid
	Phyllophaga (Chlaenobia) Blanchard, 1851 -- valid
	Phyllophaga (Cnemarachis) Saylor, 1942 -- valid
	Phyllophaga (Listrochelus) Blanchard, 1851 -- valid
	Phyllophaga (Phyllophaga) Harris, 1827 -- valid
	Phyllophaga (Phytalus) Erichson, 1847 -- valid
	Phyllophaga (Tostegoptera) Blanchard, 1851 -- valid
	Phyllophaga (Triodonyx) Saylor, 1942 -- valid
	Phyllophaga aemula (Horn, 1887) -- valid
	Phyllophaga aequalis (LeConte, 1854) -- valid
	Phyllophaga affabilis (Horn, 1887) -- valid
	Phyllophaga albina (Burmeister, 1855) -- valid
	Phyllophaga almada Saylor, 1941 -- valid
	Phyllophaga angulata Glasgow, 1925 -- invalid
	Phyllophaga anomaloides (Bates, 1888) -- valid
	Phyllophaga antennata (Smith, 1889) -- valid
	Phyllophaga anxia (LeConte, 1850) -- valid
	Phyllophaga apicata Reinhard, 1939 -- valid
	Phyllophaga arcta (Horn, 1887) -- valid
	Phyllophaga arenicola Howden, 1960 -- valid
	Phyllophaga arizona Saylor, 1940 -- valid
	Phyllophaga arizonae von Bloeker, 1936 -- invalid
	Phyllophaga arkansana (Schaeffer, 1906) -- valid
	Phyllophaga arrowi Chapin, 1935 -- invalid
	Phyllophaga aurea Luginbill and Painter, 1941 -- valid
	Phyllophaga austricolia Fall, 1929 -- invalid
	Phyllophaga babicora Mor�n, 2004 -- valid
	Phyllophaga bahama Saylor, 1940 -- invalid
	Phyllophaga balia (Say, 1825) -- valid
	Phyllophaga barda (Horn, 1887) -- valid
	Phyllophaga beckeri (Moser, 1921) -- valid
	Phyllophaga bellamyi Warner and Mor�n, 1992 -- valid
	Phyllophaga bilobatata Saylor, 1939 -- valid
	Phyllophaga bipartita (Horn, 1887) -- valid
	Phyllophaga blanchardi (Arrow, 1933) -- valid
	Phyllophaga blanda Sanderson, 1958 -- valid
	Phyllophaga bottimeri Reinhard, 1950 -- valid
	Phyllophaga brevidens (Bates, 1888) -- valid
	Phyllophaga bruneri Chapin, 1932 -- valid
	Phyllophaga bueta Saylor, 1940 -- valid
	Phyllophaga cahitana Mor�n, 2001 -- valid
	Phyllophaga calceata (LeConte, 1856) -- valid
	Phyllophaga calculiventris Saylor, 1935 -- valid
	Phyllophaga callosiventris (Moser, 1921) -- valid
	Phyllophaga canipolea Saylor, 1948 -- valid
	Phyllophaga carminator (Horn, 1894) -- valid
	Phyllophaga cavata (Bates, 1888) -- valid
	Phyllophaga cazieri Saylor, 1936 -- invalid
	Phyllophaga cazieri von Bloeker, 1936 -- invalid
	Phyllophaga cazieriana Saylor, 1938 -- valid
	Phyllophaga certanca Saylor, 1943 -- valid
	Phyllophaga chapini Saylor, 1940 -- valid
	Phyllophaga chippewa Saylor, 1939 -- valid
	Phyllophaga ciudadensis (Bates, 1888) -- valid
	Phyllophaga clemens (Horn, 1887) -- valid
	Phyllophaga clypeata (Horn, 1887) -- valid
	Phyllophaga cochisa Saylor, 1940 -- valid
	Phyllophaga congrua (LeConte, 1856) -- valid
	Phyllophaga contaminata Fall, 1932 -- valid
	Phyllophaga corrosa (LeConte, 1856) -- valid
	Phyllophaga crassissima (Blanchard, 1851) -- valid
	Phyllophaga crenulata (Fr�lich, 1792) -- valid
	Phyllophaga cribrosa (LeConte, 1853) -- valid
	Phyllophaga crinita (Burmeister, 1855) -- valid
	Phyllophaga cristagalli (Arrow, 1933) -- valid
	Phyllophaga cupuliformis Langston, 1924 -- valid
	Phyllophaga curialis Reinhard, 1939 -- valid
	Phyllophaga cushmani Saylor, 1940 -- valid
	Phyllophaga cylindrica (Burmeister, 1855) -- valid
	Phyllophaga davisi Langston, 1927 -- valid
	Phyllophaga deanii Luginbill, 1928 -- invalid
	Phyllophaga debilis (LeConte, 1856) -- valid
	Phyllophaga delata (Horn, 1887) -- valid
	Phyllophaga densicollis (LeConte, 1863) -- valid
	Phyllophaga dentex (Bates, 1888) -- valid
	Phyllophaga diffinis (Blanchard, 1851) -- valid
	Phyllophaga dispar (Burmeister, 1855) -- valid
	Phyllophaga disparilis (Horn, 1878) -- valid
	Phyllophaga dissimilis Chapin, 1935 -- invalid
	Phyllophaga divertens (Bates, 1888) -- valid
	Phyllophaga drakii (Kirby, 1837) -- valid
	Phyllophaga duncani (Barrett, 1933) -- valid
	Phyllophaga durango Saylor, 1940 -- valid
	Phyllophaga durangoana (Moser, 1921) -- valid
	Phyllophaga durangosa Saylor, 1943 -- valid
	Phyllophaga duvala Robinson, 1938 -- invalid
	Phyllophaga ecostata (Horn, 1887) -- valid
	Phyllophaga eligia Sanderson, 1958 -- valid
	Phyllophaga elizoria Saylor, 1937 -- valid
	Phyllophaga elongata (Linell, 1896) -- valid
	Phyllophaga emusa Saylor, 1937 -- invalid
	Phyllophaga eniba Saylor, 1943 -- valid
	Phyllophaga ephilida (Say, 1825) -- valid
	Phyllophaga ephilida ephilida (Say, 1825) -- valid
	Phyllophaga ephilida virilis Reinhard, 1939 -- valid
	Phyllophaga epigaea (Wickham, 1903) -- valid
	Phyllophaga errans (LeConte, 1860) -- valid
	Phyllophaga estacea Saylor, 1943 -- valid
	Phyllophaga extranea Fall, 1932 -- invalid
	Phyllophaga falsa (LeConte, 1856) -- valid
	Phyllophaga falta (LeConte, 1856) -- valid
	Phyllophaga farcta (LeConte, 1856) -- valid
	Phyllophaga fervida (Fabricius, 1775) -- valid
	Phyllophaga fimbripes (LeConte, 1856) -- valid
	Phyllophaga flavipennis (Horn, 1885) -- valid
	Phyllophaga floridana Robinson, 1938 -- valid
	Phyllophaga foralita Saylor, 1938 -- valid
	Phyllophaga forbesi Glasgow, 1916 -- valid
	Phyllophaga forsteri (Burmeister, 1855) -- valid
	Phyllophaga foxii Davis, 1920 -- valid
	Phyllophaga fraterna Harris, 1842 -- valid
	Phyllophaga fraterna fraterna Harris, 1842 -- valid
	Phyllophaga fraterna mississippiensis Davis, 1920 -- valid
	Phyllophaga fucata (Horn, 1887) -- valid
	Phyllophaga fulviventris (Moser, 1918) -- valid
	Phyllophaga fusca (Fr�lich, 1792) -- valid
	Phyllophaga futilis (LeConte, 1850) -- valid
	Phyllophaga gaigei Sanderson, 1948 -- valid
	Phyllophaga galeanae Saylor, 1943 -- valid
	Phyllophaga gentryi (Saylor, 1936) -- valid
	Phyllophaga georgiana (Horn, 1885) -- valid
	Phyllophaga ghentata (Saylor, 1937) -- valid
	Phyllophaga gigantissima Saylor, 1935 -- valid
	Phyllophaga glaberrima (Blanchard, 1851) -- valid
	Phyllophaga glabricula (LeConte, 1856) -- valid
	Phyllophaga gracilis (Burmeister, 1855) -- valid
	Phyllophaga granti Saylor, 1940 -- valid
	Phyllophaga gravidula (Moser, 1921) -- valid
	Phyllophaga grisiana von Bloeker, 1936 -- invalid
	Phyllophaga hamata (Horn, 1887) -- valid
	Phyllophaga hirsuta (Knoch, 1801) -- valid
	Phyllophaga hirticula (Knoch, 1801) -- valid
	Phyllophaga hirticula comosa Davis, 1920 -- valid
	Phyllophaga hirticula hirticula (Knoch, 1801) -- valid
	Phyllophaga hirtiventris (Horn, 1887) -- valid
	Phyllophaga hornii (Smith, 1889) -- valid
	Phyllophaga howei Sanderson, 1937 -- invalid
	Phyllophaga huachuca Saylor, 1940 -- valid
	Phyllophaga hubbelli Cartwright, 1946 -- valid
	Phyllophaga idonea Sanderson, 1948 -- valid
	Phyllophaga ignava (Horn, 1887) -- valid
	Phyllophaga ilicis (Knoch, 1801) -- valid
	Phyllophaga impar Davis, 1920 -- valid
	Phyllophaga implicita (Horn, 1887) -- valid
	Phyllophaga incuria Sanderson, 1942 -- valid
	Phyllophaga inepta (Horn, 1887) -- valid
	Phyllophaga infidelis (Horn, 1887) -- valid
	Phyllophaga inflexa Barrett, 1935 -- valid
	Phyllophaga inopia Sanderson, 1942 -- invalid
	Phyllophaga instabilis Blackwelder, 1944 -- valid
	Phyllophaga inversa (Horn, 1887) -- valid
	Phyllophaga invisa Riley and Wolfe, 1997 -- valid
	Phyllophaga iroides Fall, 1929 -- invalid
	Phyllophaga jonesi Sanderson, 1939 -- invalid
	Phyllophaga juvenilis (Fall, 1932) -- valid
	Phyllophaga karlsioei (Linell, 1898) -- valid
	Phyllophaga kentuckiana Ritcher, 1937 -- valid
	Phyllophaga knausii (Schaeffer, 1907) -- valid
	Phyllophaga knochii (Sch�nherr and Gyllenhal, 1817) -- valid
	Phyllophaga koehleriana Saylor, 1940 -- valid
	Phyllophaga lalanza Saylor, 1941 -- valid
	Phyllophaga lanceolata (Say, 1824) -- valid
	Phyllophaga laportaei (Blanchard, 1851) -- valid
	Phyllophaga latidens (Schaeffer, 1906) -- valid
	Phyllophaga latifrons (LeConte, 1856) -- valid
	Phyllophaga lenis (Horn, 1887) -- valid
	Phyllophaga linelli Saylor, 1937 -- invalid
	Phyllophaga linsleyi Saylor, 1936 -- invalid
	Phyllophaga lobata (Fall, 1908) -- valid
	Phyllophaga lodingi Sanderson, 1939 -- valid
	Phyllophaga longispina (Smith, 1889) -- valid
	Phyllophaga longitarsa (Say, 1824) -- valid
	Phyllophaga lota Luginbill, 1928 -- valid
	Phyllophaga luctuosa (Horn, 1887) -- valid
	Phyllophaga luginbilli Saylor, 1941 -- valid
	Phyllophaga macgregori Mor�n, 2004 -- valid
	Phyllophaga macmurryi Saylor, 1940 -- valid
	Phyllophaga maculicollis (LeConte, 1863) -- valid
	Phyllophaga magnicornis (Moser, 1921) -- valid
	Phyllophaga marginalis (LeConte, 1856) -- valid
	Phyllophaga marginalis insolita Cartwright, 1944 -- valid
	Phyllophaga marginalis marginalis (LeConte, 1856) -- valid
	Phyllophaga mariana Fall, 1929 -- valid
	Phyllophaga meadei Saylor, 1940 -- valid
	Phyllophaga mecocera Blackwelder, 1944 -- invalid
	Phyllophaga micans (Knoch, 1801) -- valid
	Phyllophaga michelbacheri Saylor, 1940 -- valid
	Phyllophaga microcera (Arrow, 1933) -- valid
	Phyllophaga microdon Fall, 1929 -- invalid
	Phyllophaga micros (Bates, 1888) -- valid
	Phyllophaga mimicana Saylor, 1938 -- valid
	Phyllophaga miraflora Saylor, 1940 -- valid
	Phyllophaga misteca (Bates, 1888) -- valid
	Phyllophaga molopia (Bates, 1888) -- valid
	Phyllophaga monstrosa (Saylor, 1935) -- valid
	Phyllophaga mucorea (LeConte, 1856) -- valid
	Phyllophaga murrea Sanderson, 1950 -- invalid
	Phyllophaga nebulosa Polihronakis, 2007 -- valid
	Phyllophaga neomexicana Saylor, 1940 -- valid
	Phyllophaga nitida (LeConte, 1856) -- valid
	Phyllophaga nogales Saylor, 1940 -- valid
	Phyllophaga nogueirana Mor�n, 2002 -- valid
	Phyllophaga nosa Blackwelder, 1944 -- valid
	Phyllophaga obsoleta (Blanchard, 1851) -- valid
	Phyllophaga okeechobea Robinson, 1948 -- valid
	Phyllophaga omani Sanderson, 1937 -- valid
	Phyllophaga opaca (Moser, 1918) -- valid
	Phyllophaga opacicollis (Horn, 1878) -- valid
	Phyllophaga opacita Reinhard, 1939 -- invalid
	Phyllophaga ovalis (Cartwright, 1939) -- valid
	Phyllophaga pagilis Saylor, 1937 -- invalid
	Phyllophaga pallida (Horn, 1885) -- valid
	Phyllophaga panorpa Sanderson, 1950 -- valid
	Phyllophaga parilis (Bates, 1888) -- valid
	Phyllophaga parvidens (LeConte, 1856) -- valid
	Phyllophaga parvidens hysteropyga Davis, 1920 -- valid
	Phyllophaga parvidens parvidens (LeConte, 1856) -- valid
	Phyllophaga paternoi Glasgow, 1925 -- valid
	Phyllophaga pearliae Davis, 1920 -- valid
	Phyllophaga peninsularis Saylor, 1940 -- valid
	Phyllophaga pentaphylla (Bates, 1888) -- valid
	Phyllophaga perita Sanderson, 1937 -- invalid
	Phyllophaga perlonga Davis, 1920 -- valid
	Phyllophaga pilosipes Saylor, 1940 -- valid
	Phyllophaga pilositarsis Blackwelder, 1944 -- valid
	Phyllophaga planeta Reinhard, 1950 -- valid
	Phyllophaga plena (Fall, 1932) -- valid
	Phyllophaga plena Reinhard, 1939 -- invalid
	Phyllophaga pleroma Reinhard, 1940 -- valid
	Phyllophaga ponerosa Blackwelder, 1944 -- invalid
	Phyllophaga postrema (Horn, 1887) -- valid
	Phyllophaga praesidii (Bates, 1888) -- valid
	Phyllophaga praetermissa (Horn, 1887) -- valid
	Phyllophaga profunda (Blanchard, 1851) -- valid
	Phyllophaga pruinosa (Blanchard, 1851) -- valid
	Phyllophaga prunina (LeConte, 1856) -- valid
	Phyllophaga prununculina (Burmeister, 1855) -- valid
	Phyllophaga pseudofloridana Woodruff and Beck, 1989 -- valid
	Phyllophaga psiloptera Sanderson, 1939 -- valid
	Phyllophaga pudorosa Reinhard, 1939 -- valid
	Phyllophaga pulcher (Linell, 1896) -- valid
	Phyllophaga pusillidens Fall, 1937 -- valid
	Phyllophaga querca (Knoch, 1801) -- valid
	Phyllophaga quetzala Mor�n, 2001 -- valid
	Phyllophaga ravida (Blanchard, 1851) -- valid
	Phyllophaga reevesi Saylor, 1939 -- valid
	Phyllophaga regiomontana Mor�n, 2001 -- valid
	Phyllophaga reinhardi Saylor, 1940 -- valid
	Phyllophaga renodis Reinhard, 1939 -- valid
	Phyllophaga riviera Reinhard, 1950 -- valid
	Phyllophaga rolbakeri Saylor, 1940 -- valid
	Phyllophaga rolstoni Riley and Wolfe, 1997 -- valid
	Phyllophaga rossi Saylor, 1939 -- valid
	Phyllophaga rubiginosa (LeConte, 1856) -- valid
	Phyllophaga rubricosa Reinhard, 1939 -- valid
	Phyllophaga rugipennis (Schaufuss, 1858) -- valid
	Phyllophaga rugosa (Melsheimer, 1845) -- valid
	Phyllophaga sacoma Reinhard, 1939 -- valid
	Phyllophaga sandersonia Saylor, 1939 -- invalid
	Phyllophaga saylori Sanderson, 1965 -- valid
	Phyllophaga schaefferi Saylor, 1937 -- valid
	Phyllophaga schenklingi Moser, 1918 -- valid
	Phyllophaga scitula (Horn, 1887) -- valid
	Phyllophaga scoparia (LeConte, 1856) -- valid
	Phyllophaga scuticeps (Bates, 1888) -- valid
	Phyllophaga segregans (Bates, 1888) -- valid
	Phyllophaga senex (Horn, 1878) -- valid
	Phyllophaga sequoiana Saylor, 1936 -- valid
	Phyllophaga seri Mor�n, 2002 -- invalid
	Phyllophaga setifera (Burmeister, 1855) -- valid
	Phyllophaga sinaloana Saylor, 1935 -- valid
	Phyllophaga sinuaticeps (Moser, 1921) -- valid
	Phyllophaga skelleyi Woodruff and Beck, 1989 -- valid
	Phyllophaga snowi Saylor, 1940 -- valid
	Phyllophaga sociata (Horn, 1878) -- valid
	Phyllophaga sodalis Reinhard, 1940 -- valid
	Phyllophaga sonora Saylor, 1939 -- valid
	Phyllophaga soror Davis, 1920 -- valid
	Phyllophaga spreta (Horn, 1887) -- valid
	Phyllophaga squamipilosa Saylor, 1937 -- valid
	Phyllophaga stohleri Saylor, 1938 -- valid
	Phyllophaga submucida (LeConte, 1856) -- valid
	Phyllophaga subpruinosa (Casey, 1884) -- valid
	Phyllophaga subtonsa (LeConte, 1856) -- valid
	Phyllophaga sylvatica Sanderson, 1942 -- valid
	Phyllophaga tarsalis (Schaeffer, 1908) -- valid
	Phyllophaga taxodii Langston, 1924 -- valid
	Phyllophaga tecta Cartwright, 1944 -- valid
	Phyllophaga temaxa Saylor, 1940 -- valid
	Phyllophaga temora Saylor, 1943 -- valid
	Phyllophaga texana Sanderson, 1942 -- invalid
	Phyllophaga texensis Saylor, 1940 -- valid
	Phyllophaga timida (Horn, 1878) -- valid
	Phyllophaga torta (LeConte, 1856) -- valid
	Phyllophaga trichodes (Bates, 1890) -- valid
	Phyllophaga tristis (Fabricius, 1781) -- valid
	Phyllophaga tristis amplicornis Reinhard, 1939 -- valid
	Phyllophaga tristis suttonana Reinhard, 1939 -- valid
	Phyllophaga tristis tristis (Fabricius, 1781) -- valid
	Phyllophaga trochanter Saylor, 1940 -- valid
	Phyllophaga tusa (Horn, 1887) -- valid
	Phyllophaga ulkei (Smith, 1889) -- valid
	Phyllophaga uniformis (Blanchard, 1851) -- valid
	Phyllophaga vehemens (Horn, 1887) -- valid
	Phyllophaga venodiola Saylor, 1948 -- valid
	Phyllophaga verruciventris (Moser, 1918) -- valid
	Phyllophaga vetula (Horn, 1887) -- valid
	Phyllophaga vexata (Horn, 1885) -- valid
	Phyllophaga vexata vexata (Horn, 1885) -- valid
	Phyllophaga vilifrons (LeConte, 1856) -- valid
	Phyllophaga wickhami Saylor, 1940 -- valid
	Phyllophaga xanthe (Bates, 1888) -- valid
	Phyllophaga xerophila Saylor, 1939 -- valid
	Phyllophaga yaqui Saylor, 1940 -- valid
	Phyllophaga yemasseei Cartwright, 1944 -- valid
	Phyllophaga youngi Cartwright, 1935 -- valid
	Phyllophaga zavalana Reinhard, 1946 -- valid
	Phyllostoma Fall, 1901 -- invalid
	Phytalus aegrotus Bates, 1888 -- invalid
	Phytalus cavifrons Linell, 1896 -- invalid
	Phytalus cephalicus Horn, 1885 -- invalid
	Phytalus debilis Horn, 1885 -- invalid
	Phytalus laevigatus Blanchard, 1851 -- invalid
	Phytalus longicornis Burmeister, 1855 -- invalid
	Phytalus oxypygus Burmeister, 1855 -- invalid
	Phytalus robustus Horn, 1885 -- invalid
	Phytalus vanalleri Schaeffer, 1927 -- invalid
	Pinotus Erichson, 1847 -- invalid
	Placopterus Chalumeau, 1983 -- invalid
	Plagiosalia Casey, 1915 -- invalid
	Planolinellus Dellacasa and Dellacasa, 2005 -- valid
	Planolinellus vittatus (Say, 1825) -- valid
	Planolinoides Dellacasa and Dellacasa, 2005 -- valid
	Planolinoides aenictus (Cooper and Gordon, 1987) -- valid
	Planolinoides borealis (Gyllenhal, 1827) -- valid
	Planolinoides duplex (LeConte, 1878) -- valid
	Planolinoides neotomae (Fall, 1907) -- valid
	Planolinoides pectoralis (LeConte, 1857) -- valid
	Planolinus Mulsant and Rey, 1870 -- valid
	Planolinus tenellus (Say, 1823) -- valid
	Platycerini Mulsant, 1842 -- valid
	Platyceroides Benesh, 1946 -- valid
	Platyceroides aeneus (Van Dyke, 1928) -- valid
	Platyceroides agassii (LeConte, 1861) -- valid
	Platyceroides laticollis (Casey, 1914) -- valid
	Platyceroides latus (Fall, 1901) -- valid
	Platyceroides opacus (Fall, 1906) -- valid
	Platyceroides thoracicus (Casey, 1895) -- valid
	Platyceroides viriditinctus (Benesh, 1942) -- valid
	Platyceroidini Paulsen and Hawks, 2008 -- valid
	Platyceropsis Benesh, 1946 -- valid
	Platyceropsis keeni (Casey, 1895) -- valid
	Platycerus Geoffroy, 1762 -- valid
	Platycerus angustus Casey, 1909 -- invalid
	Platycerus balbi Laporte, 1840 -- invalid
	Platycerus californicus Casey, 1885 -- invalid
	Platycerus chalybaeus Casey, 1897 -- invalid
	Platycerus coerulescens LeConte, 1869 -- invalid
	Platycerus cribripennis Van Dyke, 1928 -- invalid
	Platycerus depressus LeConte, 1850 -- valid
	Platycerus iowanus Casey, 1909 -- invalid
	Platycerus marginalis Casey, 1897 -- valid
	Platycerus oregonensis Westwood, 1846 -- valid
	Platycerus pacificus Casey, 1889 -- invalid
	Platycerus parvicollis Casey, 1889 -- invalid
	Platycerus pedicellaris M�llenkamp, 1911 -- invalid
	Platycerus peregrinus Casey, 1909 -- invalid
	Platycerus pygmaeus Van Dyke, 1946 -- invalid
	Platycerus securidens Say, 1824 -- invalid
	Platycerus virescens (Fabricius, 1775) -- valid
	Platytomus Mulsant, 1842 -- valid
	Platytomus atlanticus (Cartwright, 1948) -- valid
	Platytomus caelicollis (Cartwright, 1948) -- valid
	Platytomus longulus (Cartwright, 1948) -- valid
	Platytomus micros (Bates, 1887) -- valid
	Platytomus notialis (Cartwright, 1948) -- valid
	Platytomus parvulus (Chevrolat, 1864) -- valid
	Platytomus tibialis (Fabricius, 1798) -- valid
	Plectris LePeletier and Serville, 1828 -- valid
	Plectris aliena Chapin, 1934 -- valid
	Plectrodes Horn, 1867 -- valid
	Plectrodes pubescens Horn, 1867 -- valid
	Pleocoma LeConte, 1856 -- valid
	Pleocoma adjuvans Crotch, 1873 -- invalid
	Pleocoma australis Fall, 1911 -- valid
	Pleocoma badia Fall, 1917 -- valid
	Pleocoma badia badia Fall, 1917 -- valid
	Pleocoma badia hirsuta Davis, 1934 -- valid
	Pleocoma behrensii LeConte, 1874 -- valid
	Pleocoma bicolor Linsley, 1935 -- valid
	Pleocoma blaisdelli Linsley, 1938 -- valid
	Pleocoma carinata Linsley, 1938 -- valid
	Pleocoma conjungens Horn, 1888 -- valid
	Pleocoma conjungens conjungens Horn, 1888 -- valid -- Santa Cruz rain beetle
	Pleocoma conjungens lucia Linsley, 1941 -- valid
	Pleocoma crinita Linsley, 1938 -- valid
	Pleocoma dubitabilis Davis, 1935 -- valid
	Pleocoma dubitabilis dubitabilis Davis, 1935 -- valid
	Pleocoma dubitabilis leachi Linsley, 1938 -- valid
	Pleocoma edwardsii LeConte, 1874 -- invalid
	Pleocoma fimbriata LeConte, 1856 -- valid
	Pleocoma hirticollis Schaufuss, 1870 -- valid
	Pleocoma hirticollis hirticollis Schaufuss, 1870 -- valid
	Pleocoma hirticollis reflexa Hovore, 1972 -- valid
	Pleocoma hirticollis vandykei Linsley, 1938 -- valid
	Pleocoma hoppingi Fall, 1906 -- valid
	Pleocoma hovorei La Rue, 2007 -- valid
	Pleocoma linsleyi Hovore, 1971 -- valid
	Pleocoma marquai Hovore, 1972 -- valid
	Pleocoma minor Linsley, 1938 -- valid
	Pleocoma nitida Linsley, 1941 -- invalid
	Pleocoma octopagina Robertson, 1970 -- valid
	Pleocoma oregonensis Leach, 1933 -- valid
	Pleocoma puncticollis Rivers, 1889 -- valid
	Pleocoma remota Davis, 1934 -- invalid
	Pleocoma rickseckeri Horn, 1888 -- valid
	Pleocoma rubiginosa Hovore, 1972 -- valid
	Pleocoma rubiginosa rubiginosa Hovore, 1972 -- valid
	Pleocoma rubiginosa transsierrae Hovore, 1972 -- valid
	Pleocoma shastensis Van Dyke, 1933 -- valid
	Pleocoma simi Davis, 1934 -- valid
	Pleocoma sonomae Linsley, 1935 -- valid
	Pleocoma staff Schaufuss, 1870 -- valid
	Pleocoma trifoliata Linsley, 1938 -- valid
	Pleocoma tularensis Leach, 1933 -- valid
	Pleocoma ulkei Horn, 1888 -- invalid
	Pleocoma venturae Linsley, 1945 -- invalid
	Pleocomidae LeConte, 1861 -- valid
	Pleopertha Reitter, 1903 -- invalid
	Plesiosternus Mor�n, 1983 -- valid
	Plesiosternus setosus Mor�n, 1983 -- valid
	Pleurophorus Mulsant, 1842 -- valid
	Pleurophorus batesi Arrow, 1903 -- invalid
	Pleurophorus caesus (Panzer, 1796) -- valid
	Plusiotina Casey, 1915 -- invalid
	Plusiotina aeruginis Casey, 1915 -- invalid
	Plusiotina angusta Casey, 1915 -- invalid
	Plusiotina sonorica Casey, 1915 -- invalid
	Plusiotina subenodis Casey, 1915 -- invalid
	Plusiotis Burmeister, 1844 -- invalid
	Plusiotis ampliata Casey, 1915 -- invalid
	Plusiotis ocularis Casey, 1915 -- invalid
	Pocalta Casey, 1915 -- invalid
	Pocalta brevis Casey, 1915 -- invalid
	Pocalta laevicauda Casey, 1915 -- invalid
	Pocalta nigripennis Casey, 1915 -- invalid
	Pocalta pubicollis Casey, 1915 -- invalid
	Pocalta rotunda Casey, 1915 -- invalid
	Pocalta rubripennis Casey, 1915 -- invalid
	Pocalta seriata Casey, 1915 -- invalid
	Podalgus obesus Burmeister, 1847 -- invalid
	Podalgus variolosus Burmeister, 1847 -- invalid
	Podolasia Harold, 1869 -- valid
	Podolasia emarginata Howden, 1954 -- valid
	Podolasia ferruginea (LeConte, 1856) -- valid
	Podolasia involucris Howden, 1998 -- valid
	Podolasia lavignei Howden, 1997 -- valid
	Podolasia longipennis Howden, 1997 -- valid
	Podolasia peninsularis Howden, 1954 -- valid
	Podolasia pilosa Howden, 1954 -- valid
	Podolasia rotundipenis Howden, 1997 -- valid
	Podolasia saylori Howden, 1954 -- invalid
	Podolasia stillwellorum Howden, 1997 -- valid
	Podolasia varicolor Saylor, 1948 -- valid
	Podolasiini Howden, 1997 -- valid
	Podostena Howden, 1997 -- valid
	Podostena bottimeri (Howden, 1958) -- valid
	Podostena litoralis Howden, 1997 -- valid
	Podostena rileyi Howden, 1997 -- valid
	Podostena sleeperi Howden, 1997 -- valid
	Polyacanthopus Kuwert, 1896 -- invalid
	Polylamina Hardy, 1974 -- invalid
	Polylamina pubescens (Cartwright, 1939) -- invalid -- Wooly Gulf dune scarab
	Polymoechus LeConte, 1856 -- invalid
	Polymoechus conicicollis Casey, 1915 -- invalid
	Polymoechus discernens Casey, 1915 -- invalid
	Polyphylla Harris, 1841 -- valid
	Polyphylla (Polyphylla) Harris, 1841 -- valid
	Polyphylla acomana Casey, 1914 -- invalid
	Polyphylla adusta Casey, 1914 -- invalid
	Polyphylla aeola La Rue, 1998 -- valid
	Polyphylla alleni Cazier, 1939 -- invalid
	Polyphylla anteronivea Hardy, 1978 -- valid -- Saline Valley snow-front June beetle
	Polyphylla arguta Casey, 1914 -- valid
	Polyphylla arida Van Dyke, 1947 -- invalid
	Polyphylla avittata Hardy, 1978 -- valid -- Spotted Warner Valley Dunes June beetle
	Polyphylla barbata Cazier, 1938 -- valid -- Mount Hermon June beetle
	Polyphylla bisinuata Casey, 1914 -- invalid
	Polyphylla brownae Young, 1986 -- valid
	Polyphylla castanea Casey, 1914 -- invalid
	Polyphylla cavifrons LeConte, 1854 -- valid
	Polyphylla comes Casey, 1914 -- valid
	Polyphylla comstockiana von Bloeker, 1939 -- invalid
	Polyphylla crinita LeConte, 1856 -- valid
	Polyphylla decemlineata (Say, 1824) -- valid
	Polyphylla devestiva Young, 1966 -- valid
	Polyphylla diffracta Casey, 1891 -- valid
	Polyphylla diffusa Casey, 1914 -- invalid
	Polyphylla donaldsoni Skelley, 2005 -- valid
	Polyphylla dubia Hatch, 1971 -- invalid
	Polyphylla erratica Hardy, 1978 -- valid -- Death Valley June beetle
	Polyphylla fuscula Fall, 1908 -- invalid
	Polyphylla gracilis Horn, 1881 -- valid
	Polyphylla hammondi LeConte, 1856 -- valid
	Polyphylla hirsuta Van Dyke, 1933 -- valid
	Polyphylla impigra Casey, 1914 -- invalid
	Polyphylla incolumis Casey, 1914 -- invalid
	Polyphylla laevicauda Casey, 1914 -- invalid
	Polyphylla laticauda Casey, 1914 -- invalid
	Polyphylla latifrons Casey, 1914 -- invalid
	Polyphylla martini von Bloeker, 1939 -- invalid
	Polyphylla matrona Casey, 1914 -- invalid
	Polyphylla mescalerensis Young, 1988 -- valid
	Polyphylla modulata Casey, 1914 -- valid
	Polyphylla molesta Casey, 1914 -- invalid
	Polyphylla monahansensis Hardy, 1978 -- valid
	Polyphylla multimaculata Hardy, 1981 -- valid
	Polyphylla mystica Casey, 1914 -- invalid
	Polyphylla nigra Casey, 1914 -- valid
	Polyphylla nubila Van Dyke, 1947 -- valid -- Atascodero June beetle
	Polyphylla oblita Casey, 1914 -- invalid
	Polyphylla occidentalis (Linnaeus, 1767) -- valid
	Polyphylla oklahomensis Hatch, 1926 -- invalid
	Polyphylla ona von Bloeker, 1939 -- invalid
	Polyphylla opposita Casey, 1914 -- invalid
	Polyphylla oregona Casey, 1914 -- invalid
	Polyphylla pacifica Casey, 1895 -- invalid
	Polyphylla parilis Casey, 1914 -- invalid
	Polyphylla peninsularis Van Dyke, 1947 -- invalid
	Polyphylla perversa Casey, 1914 -- invalid
	Polyphylla petitii (Gu�rin-M�neville, 1844) -- valid
	Polyphylla pimalis Casey, 1914 -- invalid
	Polyphylla potosiana Casey, 1914 -- invalid
	Polyphylla pottsorum Hardy, 1978 -- valid
	Polyphylla proba Casey, 1914 -- invalid
	Polyphylla pubescens Cartwright, 1939 -- valid -- Wooly Gulf dune scarab
	Polyphylla ratcliffei Young, 1986 -- valid
	Polyphylla reducta Casey, 1914 -- invalid
	Polyphylla relicta Casey, 1914 -- invalid
	Polyphylla robustula Casey, 1914 -- invalid
	Polyphylla rufescenta Tanner, 1928 -- invalid
	Polyphylla ruficollis Casey, 1914 -- invalid
	Polyphylla rugosipennis Casey, 1914 -- valid
	Polyphylla santacruzae von Bloeker, 1939 -- invalid
	Polyphylla santarosae von Bloeker, 1939 -- invalid
	Polyphylla sejuncta Casey, 1914 -- invalid
	Polyphylla sobrina Casey, 1914 -- valid
	Polyphylla speciosa Casey, 1889 -- invalid
	Polyphylla squamicauda Casey, 1914 -- invalid
	Polyphylla squamiventris Cazier, 1939 -- valid
	Polyphylla squamotecta Casey, 1914 -- invalid
	Polyphylla stellata Young, 1986 -- valid -- Delta June beetle
	Polyphylla subvittata LeConte, 1856 -- invalid
	Polyphylla uteana Tanner, 1928 -- invalid
	Polyphylla variolosa (Hentz, 1830) -- valid
	Polyphylla verecunda Casey, 1914 -- invalid
	Polyphylla woodruffi Skelley, 2005 -- valid
	Popillia Dejean, 1821 -- valid
	Popillia japonica Newman, 1841 -- valid
	Popillia plicatipennis Burmeister, 1844 -- invalid
	Popilliina Ohaus, 1918 -- valid
	Proctophanini Stebnicka and Howden, 1995 -- valid
	Proculini Kaup, 1868 -- valid
	Procululus Zang, 1905 -- invalid
	Protaetia Burmeister, 1842 -- valid
	Protaetia (Heteroprotaetia) Miksic, 1963 -- valid
	Protaetia bourgoini Paulian, 1960 -- invalid
	Protaetia fusca (Herbst, 1790) -- valid
	Protaetia taiwana Nijima and Kinoshita, 1923 -- invalid
	Psammobius Heer, 1841 -- invalid
	Psammodiini Mulsant, 1842 -- valid
	Psammodius Fall�n, 1807 -- valid
	Psammodius aegialioides Haldeman, 1848 -- invalid
	Psammodius basalis (Mulsant and Rey, 1870) -- valid
	Psammodius grundeli Van Dyke, 1918 -- invalid
	Psammodius laevipennis Costa, 1844 -- valid
	Psammodius perpolitus Schmidt, 1916 -- invalid
	Psammodius pierottii Pittino, 1979 -- valid
	Psammodius schwarzi Linell, 1896 -- invalid
	Psammodius shermani Cartwright, 1946 -- invalid
	Pseudacratus Dalla Torre, 1912 -- invalid
	Pseudagolius Schmidt, 1913 -- valid
	Pseudagolius bicolor (Say, 1823) -- valid
	Pseudagolius coloradensis (Horn, 1870) -- valid
	Pseudagolius warneri Gordon and Skelley, 2007 -- valid
	Pseudammoecius Schmidt, 1913 -- invalid
	Pseudancognatha Otoya, 1945 -- invalid
	Pseudaphonus Casey, 1915 -- invalid
	Pseudaphonus debiliceps Casey, 1915 -- invalid
	Pseudaphonus lucidus Casey, 1915 -- invalid
	Pseudaphonus ovalis Casey, 1915 -- invalid
	Pseudaphonus puncticollis Casey, 1924 -- invalid
	Pseudaphonus repens Casey, 1915 -- invalid
	Pseudataenius Brown, 1927 -- valid
	Pseudataenius contortus Cartwright, 1974 -- valid
	Pseudataenius socialis (Horn, 1871) -- valid
	Pseudocalamosternus Balthasar, 1936 -- invalid
	Pseudocanthon Bates, 1887 -- valid
	Pseudocanthon perplexus (LeConte, 1847) -- valid
	Pseudochodaeini Scholtz in Scholtz, D'hotman, Evans and Nel, 1988 -- valid
	Pseudochodaeus Carlson and Ritcher, 1974 -- valid
	Pseudochodaeus estriatus (Schaeffer, 1906) -- valid
	Pseudocotalpa Hardy, 1971 -- valid
	Pseudocotalpa andrewsi Hardy, 1971 -- valid -- Andrews' dune scarab beetle
	Pseudocotalpa giulianii Hardy, 1974 -- valid -- Giuliani's dune scarab
	Pseudocotalpa sonorica Hardy, 1974 -- valid
	Pseudolucanus Hope, 1845 -- invalid
	Pseudolucanus bicostatus Angell, 1917 -- invalid
	Pseudolucanus capreolus (Linnaeus, 1764) -- invalid -- stag beetle
	Pseudomaladera Nikolajev, 1980 -- invalid
	Pseudopopillia Kolbe, 1894 -- invalid
	Pseudoserica Gu�rin-M�neville, 1838 -- invalid
	Pseudotrox Robinson, 1948 -- invalid
	Psilax Erichson, 1848 -- invalid
	Psilocnemis Burmeister, 1842 -- valid
	Psilocnemis leucosticta Burmeister, 1842 -- valid
	Psilomus Zang, 1905 -- invalid
	Ptichopus Kaup, 1869 -- valid
	Ptichopus aberrator Kuwert, 1891 -- invalid
	Ptichopus angulatus (Percheron, 1835) -- valid
	Ptichopus borellii Rosmini, 1902 -- invalid
	Ptichopus inca Kuwert, 1891 -- invalid
	Ptichopus inflatus Kuwert, 1898 -- invalid
	Ptichopus montezuma Kuwert, 1891 -- invalid
	Ptichopus nitidus Kuwert, 1891 -- invalid
	Ptychotrichus Kuwert, 1896 -- invalid
	Rhagonocerus Kaup, 1871 -- invalid
	Rhipiphoria Casey, 1915 -- invalid
	Rhizotrogus collocatus Walker, 1866 -- invalid
	Rhodocanthopus Kaup, 1871 -- invalid
	Rhombonalia Casey, 1915 -- invalid
	Rhombonalia cochiseana Casey, 1915 -- invalid
	Rhombonalia comes Casey, 1915 -- invalid
	Rhombonalia transversa Casey, 1915 -- invalid
	Rhysothorax Bedel, 1911 -- valid
	Rhysothorax rufus (Fabricius, 1792) -- valid
	Rhyssemus Mulsant, 1842 -- valid
	Rhyssemus aspericeps Chevrolat, 1861 -- invalid
	Rhyssemus brownwoodi Gordon and Cartwright, 1980 -- valid
	Rhyssemus caelatus LeConte, 1881 -- invalid
	Rhyssemus californicus Horn, 1871 -- valid
	Rhyssemus cribrosus LeConte, 1850 -- invalid
	Rhyssemus germanus (Linnaeus, 1767) -- valid
	Rhyssemus neglectus Brown, 1929 -- valid
	Rhyssemus obsoletus Rey, 1890 -- invalid
	Rhyssemus parallelus Reitter, 1892 -- invalid
	Rhyssemus puncticollis Brown, 1929 -- invalid
	Rhyssemus scaber Haldeman, 1848 -- valid
	Rhyssemus sonatus LeConte, 1881 -- valid
	Roplisa Casey, 1909 -- invalid
	Rugaphodius Gordon and Skelley, 2007 -- valid
	Rugaphodius rugatus (Schmidt, 1907) -- valid
	Rutela Latreille, 1802 -- valid
	Rutela formosa Burmeister, 1844 -- valid
	Rutelinae MacLeay, 1819 -- valid
	Rutelini MacLeay, 1819 -- valid
	Rutelisca Bates, 1888 -- valid
	Rutelisca durangoana Ohaus, 1905 -- valid
	Saprosites nocturnus Nomura, 1943 -- invalid
	Saprosites rugosus Richards, 1959 -- invalid
	Sayloria Chalumeau, 1981 -- invalid
	Scabrostomus Gordon and Skelley, 2007 -- valid
	Scabrostomus baileyi (Skelley and Gordon, 2002) -- valid
	Scabrostomus bakeri (Skelley and Gordon, 2002) -- valid
	Scabrostomus dyspistus (Skelley and Woodruff, 1991) -- valid
	Scabrostomus kerni Gordon and Skelley, 2007 -- valid
	Scabrostomus parapeculiosus Gordon and Skelley, 2007 -- valid
	Scabrostomus peculiosus (Schmidt, 1922) -- valid
	Scabrostomus sepultus (Cartwright, 1944) -- valid
	Scabrostomus siccatus Gordon and Skelley, 2007 -- valid
	Scabrostomus skillmani (Warner and Skelley, 2006) -- valid
	Scabrostomus tanytarsus (Skelley and Woodruff, 1991) -- valid
	Scalmus Zang, 1905 -- invalid
	Scarabaeidae Latreille, 1802 -- valid -- scarab beetles
	Scarabaeinae Latreille, 1802 -- valid
	Scarabaeoidea Latreille, 1802 -- valid
	Scarabaeus americanus Palisot de Beauvois, 1807 -- invalid
	Scarabaeus ammon Fabricius, 1781 -- invalid
	Scarabaeus antillarum Palisot de Beauvois, 1811 -- invalid
	Scarabaeus arator Illiger, 1803 -- invalid
	Scarabaeus asper Fabricius, 1775 -- invalid
	Scarabaeus attaminatus Marsham, 1802 -- invalid
	Scarabaeus autumnalis Naezen, 1792 -- invalid
	Scarabaeus bicolor Fourcroy, 1785 -- invalid
	Scarabaeus biliteratus Marsham, 1802 -- invalid
	Scarabaeus boscii Palisot de Beauvois, 1809 -- invalid
	Scarabaeus capitatus De Geer, 1774 -- invalid
	Scarabaeus capra Fabricius, 1787 -- invalid
	Scarabaeus centrolineatus Panzer, 1798 -- invalid
	Scarabaeus cephus Fabricius, 1775 -- invalid
	Scarabaeus cruoreus Schrank, 1798 -- invalid
	Scarabaeus dorcas Olivier, 1789 -- invalid
	Scarabaeus fenestralis Schrank, 1781 -- invalid
	Scarabaeus flagranticeps Voet, 1766 -- invalid
	Scarabaeus foedatus Marsham, 1802 -- invalid
	Scarabaeus foveolatus Moll, 1785 -- invalid
	Scarabaeus fuscus Rossi, 1792 -- invalid
	Scarabaeus globosus Kugelann, 1794 -- invalid
	Scarabaeus hudsonias Forster, 1771 -- invalid
	Scarabaeus inquinatus Herbst, 1783 -- invalid
	Scarabaeus janus Panzer, 1794 -- invalid
	Scarabaeus juvencus Fabricius, 1775 -- invalid
	Scarabaeus laevis Drury, 1773 -- invalid
	Scarabaeus lar Fabricius, 1787 -- invalid
	Scarabaeus latipes Germar, 1824 -- invalid
	Scarabaeus maimon Fabricius, 1775 -- invalid
	Scarabaeus marianus Linnaeus, 1767 -- invalid
	Scarabaeus meliboeus Fabricius, 1792 -- invalid
	Scarabaeus minutus Herbst, 1789 -- invalid
	Scarabaeus nubilus Panzer, 1798 -- invalid
	Scarabaeus oblongus Scopoli, 1763 -- invalid
	Scarabaeus pedellus De Geer, 1774 -- invalid
	Scarabaeus pennsylvanicus De Geer, 1774 -- invalid
	Scarabaeus porcatus Fabricius, 1775 -- invalid
	Scarabaeus quadrituberculatus Fabricius, 1798 -- invalid
	Scarabaeus quadrum Kugelann, 1792 -- invalid
	Scarabaeus recticornis Leske, 1785 -- invalid
	Scarabaeus rugosus Poda von Neuhaus, 1761 -- invalid
	Scarabaeus satyrus Fabricius, 1775 -- invalid
	Scarabaeus silenus Fabricius, 1775 -- invalid
	Scarabaeus suturalis Fabricius, 1792 -- invalid
	Scarabaeus tomentosus Kugelann, 1792 -- invalid
	Scarabaeus triangulus Piller and Mitterpacher, 1783 -- invalid
	Scarabaeus tullius Olivier, 1789 -- invalid
	Scarabaeus turchesinus Voet, 1778 -- invalid
	Scarabaeus variegatus Scopoli, 1763 -- invalid
	Scarabaeus volvens Fabricius, 1792 -- invalid
	Schaefferellus Gordon and Skelley, 2007 -- valid
	Schaefferellus arizonensis (Schaeffer, 1907) -- valid
	Serica MacLeay, 1819 -- valid
	Serica abdita Dawson, 1921 -- valid
	Serica acicula Dawson, 1932 -- valid
	Serica acontia Dawson, 1933 -- valid
	Serica adversa Dawson, 1967 -- valid
	Serica aemula Dawson, 1947 -- valid
	Serica alabama Dawson, 1967 -- valid
	Serica alleni Saylor, 1939 -- valid
	Serica alternata LeConte, 1856 -- valid
	Serica alternata alternata LeConte, 1856 -- valid
	Serica alternata exolita Dawson, 1933 -- valid
	Serica alternata patruela Dawson, 1933 -- valid
	Serica ammomenisco Hardy, 1987 -- valid -- Crescent Dunes serican scarab
	Serica anthracina LeConte, 1856 -- valid
	Serica apatela Dawson, 1922 -- valid
	Serica arkansana Dawson, 1947 -- valid
	Serica aspera Dawson, 1922 -- valid
	Serica atracapilla (Kirby, 1837) -- valid
	Serica atratula LeConte, 1856 -- valid
	Serica atratula atratula LeConte, 1856 -- valid
	Serica atratula monita Dawson, 1947 -- valid
	Serica aviceps Dawson, 1967 -- valid
	Serica barri Dawson, 1967 -- valid
	Serica blatchleyi Dawson, 1932 -- valid
	Serica bruneri Dawson, 1967 -- valid
	Serica caliginosa Dawson, 1932 -- valid
	Serica campestris Dawson, 1919 -- valid
	Serica carinata Blatchley, 1910 -- invalid
	Serica carolina Dawson, 1920 -- valid
	Serica catalina Dawson, 1947 -- valid
	Serica chaetosoma Dawson, 1932 -- valid
	Serica chicoensis Saylor, 1939 -- valid
	Serica coalinga Dawson, 1952 -- valid
	Serica concinna Dawson, 1947 -- valid
	Serica contorta Dawson, 1947 -- valid
	Serica craighead Saylor, 1939 -- valid
	Serica craigheadi Dawson, 1952 -- invalid
	Serica crassata Walker, 1866 -- invalid
	Serica cruzi Saylor, 1939 -- valid
	Serica cucullata Dawson, 1947 -- invalid
	Serica curvata LeConte, 1856 -- valid
	Serica cuyamaca Saylor, 1939 -- valid
	Serica delicata Dawson, 1922 -- valid
	Serica deserticola Dawson, 1952 -- valid
	Serica diablo Dawson, 1967 -- valid
	Serica egregia Dawson, 1921 -- valid
	Serica elmontea Saylor, 1939 -- valid
	Serica elongatula Horn, 1870 -- valid
	Serica elusa Dawson, 1919 -- valid
	Serica ensenada Saylor, 1948 -- valid
	Serica errans Blatchley, 1929 -- invalid
	Serica evidens Blatchley, 1919 -- valid
	Serica falcata Dawson, 1933 -- valid
	Serica falli Dawson, 1932 -- valid
	Serica fimbriata LeConte, 1856 -- valid
	Serica floridana Dawson, 1967 -- valid
	Serica frontalis LeConte, 1856 -- invalid
	Serica frosti Dawson, 1967 -- valid -- Frost's spring serican scarab
	Serica georgiana Leng, 1911 -- valid
	Serica georgiana georgiana Leng, 1911 -- valid
	Serica georgiana lecontei Dawson, 1921 -- valid
	Serica heteracantha Dawson, 1967 -- valid
	Serica howdeni Dawson, 1967 -- valid
	Serica humboldti Gordon, 1975 -- valid
	Serica imitans Chapin, 1931 -- valid
	Serica intermixta Blatchley, 1910 -- valid
	Serica iricolor (Say, 1824) -- valid
	Serica joaquinella Saylor, 1939 -- invalid
	Serica korgei Petrovitz, 1967 -- invalid
	Serica laguna Saylor, 1935 -- valid
	Serica ligulata Dawson, 1932 -- valid
	Serica ligulata ligulata Dawson, 1932 -- valid
	Serica ligulata praetermissa Dawson, 1932 -- valid
	Serica lodingi Dawson, 1952 -- valid
	Serica loxia Dawson, 1920 -- valid
	Serica mckenziei Saylor, 1935 -- valid
	Serica mendota Saylor, 1939 -- invalid
	Serica michelbacheri Saylor, 1948 -- invalid
	Serica mixta LeConte, 1856 -- valid
	Serica mystaca Dawson, 1922 -- valid
	Serica ochrosoma Dawson, 1919 -- valid
	Serica oliver Saylor, 1939 -- valid
	Serica oliveri Dawson, 1952 -- invalid
	Serica opposita Dawson, 1921 -- valid
	Serica panda Dawson, 1952 -- valid
	Serica parallela Casey, 1884 -- valid
	Serica parvula (Blanchard, 1850) -- valid
	Serica pavonia Dawson, 1932 -- valid
	Serica peleca Dawson, 1952 -- valid
	Serica perigonia Dawson, 1920 -- valid
	Serica perigonia eremicola Dawson, 1967 -- valid
	Serica perigonia perigonia Dawson, 1920 -- valid
	Serica pilifera Horn, 1894 -- valid
	Serica porcula Casey, 1884 -- valid
	Serica prava Dawson, 1933 -- valid
	Serica pruinosa Saylor, 1935 -- invalid
	Serica prunipennis Saylor, 1936 -- valid
	Serica psammobuna Hardy, 1987 -- valid
	Serica psammobunus Hardy, 1987 -- invalid -- Sand Mountain serican scarab
	Serica pullata Dawson, 1967 -- valid
	Serica pusilla Dawson, 1922 -- valid
	Serica repanda Dawson, 1933 -- valid
	Serica rhypha Dawson, 1952 -- valid
	Serica robusta LeConte, 1856 -- invalid
	Serica rossi Saylor, 1948 -- valid
	Serica sandiegensis Saylor, 1939 -- valid
	Serica satrapa Dawson, 1947 -- valid
	Serica scaphia Dawson, 1952 -- valid
	Serica sculptilis Dawson, 1922 -- valid
	Serica searli Saylor, 1939 -- invalid
	Serica senta Dawson, 1933 -- valid
	Serica serensia Saylor, 1948 -- valid
	Serica sericea (Illiger, 1802) -- valid
	Serica sericeoides Dawson, 1967 -- valid
	Serica serotina LeConte, 1856 -- valid
	Serica solita Dawson, 1922 -- valid
	Serica spicula Dawson, 1921 -- valid
	Serica sponsa Dawson, 1919 -- valid
	Serica stygia Dawson, 1933 -- valid
	Serica subnisa Dawson, 1947 -- valid
	Serica tantula Dawson, 1922 -- valid -- Tantula serican scarab
	Serica texana LeConte, 1856 -- valid
	Serica tristis LeConte, 1850 -- valid
	Serica trociformis Burmeister, 1844 -- valid
	Serica valida Harold, 1869 -- invalid
	Serica ventura Dawson, 1932 -- valid
	Serica ventura dorsalis Dawson, 1952 -- valid
	Serica ventura personata Dawson, 1932 -- valid
	Serica ventura ventura Dawson, 1932 -- valid
	Serica vespertina (Gyllenhal, 1817) -- valid
	Serica vespertina accola Dawson, 1921 -- valid
	Serica vespertina vespertina (Gyllenhal, 1817) -- valid
	Serica watson Saylor, 1939 -- valid
	Serica watsoni Dawson, 1952 -- invalid
	Sericini Kirby, 1837 -- valid
	Setodius Gordon and Skelley, 2007 -- valid
	Setodius bryanti (Brown, 1929) -- valid
	Setodius diogenes Gordon and Skelley, 2007 -- valid
	Setodius edmontonus (Brown, 1929) -- valid
	Severus Kuwert, 1896 -- invalid
	Sinodendron Hellwig, 1791 -- valid
	Sinodendron rugosum Mannerheim, 1843 -- valid
	Sisyphini Mulsant, 1842 -- valid
	Sisyphus Latreille, 1807 -- valid
	Sisyphus submonticolus Howden, 1965 -- valid
	Sphaeromorphus Germar, 1843 -- invalid
	Sphaeromorphus volvox Erichson, 1843 -- invalid
	Spilosota Casey, 1915 -- invalid
	Spilosota inconspicua Casey, 1915 -- invalid
	Spilosota magister Casey, 1915 -- invalid
	Spilosota nubeculina Casey, 1915 -- invalid
	Spilosota pallidissima Casey, 1915 -- invalid
	Spilota Burmeister, 1844 -- invalid
	Spilota incolumis Casey, 1915 -- invalid
	Spilota maritima Casey, 1915 -- invalid
	Stenobronchus Schmidt, 1916 -- invalid
	Stenothorax Harris, 1827 -- invalid
	Stenotothorax Schmidt, 1913 -- valid
	Stenotothorax badipes (Melsheimer, 1845) -- valid
	Stenotothorax cadaverinus (Mannerheim, 1843) -- valid
	Stenotothorax comosus Gordon and Skelley, 2007 -- valid
	Stenotothorax cribratulus (Schmidt, 1922) -- valid
	Stenotothorax dilaticollis (Saylor, 1935) -- valid
	Stenotothorax gardneri (Gordon, 2006) -- valid
	Stenotothorax gentilis (Horn, 1887) -- valid
	Stenotothorax gravis (Fall, 1927) -- valid
	Stenotothorax lanei (Saylor, 1940) -- valid
	Stenotothorax linsleyi (Van Dyke, 1933) -- valid
	Stenotothorax martini (Van Dyke, 1928) -- valid
	Stenotothorax mcpeaki Gordon and Skelley, 2007 -- valid
	Stenotothorax micellus Gordon and Skelley, 2007 -- valid
	Stenotothorax nevadensis (Horn, 1870) -- valid
	Stenotothorax oriens Gordon and Skelley, 2007 -- valid
	Stenotothorax oviformis (Robinson, 1946) -- valid
	Stenotothorax ovipennis (Horn, 1870) -- valid
	Stenotothorax parapyriformis Gordon and Skelley, 2007 -- valid
	Stenotothorax pyriformis (Brown, 1929) -- valid
	Stenotothorax rugoclypeus (Hinton, 1934) -- valid
	Stenotothorax sparsus (LeConte, 1878) -- valid
	Stenotothorax washtucna (Robinson, 1938) -- valid
	Stenotothorax woodleyi Gordon, 2006 -- valid
	Stephanucha Burmeister, 1842 -- invalid
	Stephanucha atrata Thomson, 1878 -- invalid
	Stephanucha thoracica Casey, 1915 -- invalid
	Stigmalia Casey, 1915 -- invalid
	Stigmalia cuernavacana Casey, 1915 -- invalid
	Stigmalia deficiens Casey, 1915 -- invalid
	Stigmalia fallaciosa Casey, 1915 -- invalid
	Stigmalia histrionica Casey, 1915 -- invalid
	Stilbolemma Harris, 1827 -- invalid
	Stomphax Fischer von Waldheim, 1823 -- invalid
	Strategodes Casey, 1915 -- invalid
	Strategopsis Chapin, 1932 -- invalid
	Strategus Kirby, 1828 -- valid
	Strategus aloeus (Linnaeus, 1758) -- valid
	Strategus antaeus (Drury, 1773) -- valid
	Strategus arizonicus Schaeffer, 1915 -- invalid
	Strategus atrolucens Casey, 1915 -- invalid
	Strategus beckeri Kolbe, 1906 -- invalid
	Strategus cessus LeConte, 1866 -- valid
	Strategus craigi Ratcliffe, 1976 -- valid
	Strategus divergens Casey, 1915 -- invalid
	Strategus gaillardi Casey, 1915 -- invalid
	Strategus houstonensis Knaus, 1925 -- invalid
	Strategus howdeni Ratcliffe, 1976 -- valid
	Strategus julianus Burmeister, 1847 -- invalid
	Strategus longichomperus Ratcliffe, 1976 -- valid
	Strategus mormon Burmeister, 1847 -- valid
	Strategus pinorum Casey, 1915 -- invalid
	Strategus roosevelti Casey, 1915 -- invalid
	Strategus semistriatus Casey, 1915 -- invalid
	Strategus septentrionis Casey, 1915 -- invalid
	Strategus sinuatus Casey, 1915 -- invalid
	Strategus splendens (Palisot de Beauvois, 1809) -- valid
	Strategus tarsalis Casey, 1915 -- invalid
	Strategus temoltzin Mor�n and Nogueira, 2008 -- valid
	Strigidia Burmeister, 1844 -- invalid
	Strigoderma Burmeister, 1844 -- valid
	Strigoderma angustula Casey, 1915 -- invalid
	Strigoderma arbicola (Fabricius, 1793) -- valid
	Strigoderma aterrima Casey, 1915 -- invalid
	Strigoderma convergens Casey, 1915 -- invalid
	Strigoderma costulipennis Bates, 1888 -- valid
	Strigoderma fastuosa Burmeister, 1844 -- invalid
	Strigoderma floridana Ohaus, 1905 -- invalid
	Strigoderma imitatrix Nonfried, 1893 -- invalid
	Strigoderma impressicollis Nonfried, 1893 -- invalid
	Strigoderma irregularis Casey, 1915 -- invalid
	Strigoderma knausi (Brown, 1925) -- valid
	Strigoderma latitibia Schaeffer, 1907 -- invalid
	Strigoderma mediocris Casey, 1915 -- invalid
	Strigoderma metallica Nonfried, 1893 -- invalid
	Strigoderma morelosana Casey, 1915 -- invalid
	Strigoderma nigripes Burmeister, 1844 -- invalid
	Strigoderma nigrita Blanchard, 1851 -- invalid
	Strigoderma obesula Casey, 1915 -- invalid
	Strigoderma pimalis Casey, 1884 -- valid
	Strigoderma prolixella Casey, 1915 -- invalid
	Strigoderma protea Burmeister, 1844 -- valid
	Strigoderma puritana Casey, 1915 -- invalid
	Strigoderma pygmaea (Fabricius, 1798) -- valid
	Strigoderma quaternaria Casey, 1915 -- invalid
	Strigoderma radula Casey, 1915 -- invalid
	Strigoderma sonorica Casey, 1915 -- invalid
	Strigoderma subrutilans Casey, 1915 -- invalid
	Strigoderma sulcipennis Burmeister, 1844 -- valid
	Strigoderma teapensis Bates, 1888 -- valid
	Strigoderma testaceipennis Nonfried, 1893 -- invalid
	Strigoderma texana Casey, 1915 -- invalid
	Strigoderma vestita Burmeister, 1844 -- valid
	Strigoderma virginica Casey, 1915 -- invalid
	Strigoderma viridicollis Schaeffer, 1907 -- invalid
	Strigodermella Casey, 1915 -- invalid
	Strigodermella oklahomensis Hatch, 1930 -- invalid
	Strigodius Gordon and Skelley, 2007 -- valid
	Strigodius robinsoni (Cartwright, 1939) -- valid
	Syndesinae MacLeay, 1819 -- valid
	Synesius Kuwert, 1896 -- invalid
	Systenocerus Weise, 1883 -- invalid
	Systenus Sharp and Muir, 1912 -- invalid
	Tauronthophagus Shipp, 1895 -- invalid
	Tesarius Rakovic, 1981 -- valid
	Tesarius caelatus (LeConte, 1857) -- valid
	Tesarius doyeni (Cartwright, 1977) -- valid
	Tesarius mcclayi (Cartwright, 1955) -- valid
	Tesarius oregonensis (Cartwright, 1955) -- valid
	Tetraclipeoides Schmidt, 1913 -- valid
	Tetraclipeoides acutissimus (Gordon, 1976) -- valid
	Tetraclipeoides aemulus (Horn, 1887) -- valid
	Tetraclipeoides anomaliceps (Brown, 1929) -- valid
	Tetraclipeoides denticulatus (Haldeman, 1848) -- valid
	Tetraclipeoides dentiger (LeConte, 1858) -- valid
	Tetraclipeoides dentigerulus (Brown, 1929) -- valid
	Tetraclipeoides fimbripes (Brown, 1928) -- valid
	Tetraclipeoides formidatus (Gordon, 1976) -- valid
	Tetraclipeoides giulianii (Gordon, 1977) -- valid
	Tetraclipeoides henryi (Gordon, 1976) -- valid
	Tetraclipeoides moquinus (Fall, 1907) -- valid
	Tetraclipeoides testaceiventris (Fall, 1932) -- valid
	Tetraclipeoides texanus Gordon and Skelley, 2007 -- valid
	Tetraracus Kuwert, 1891 -- invalid
	Teuchestes Mulsant, 1842 -- valid
	Teuchestes brunneus Mulsant, 1842 -- invalid
	Teuchestes fossor (Linnaeus, 1758) -- valid
	Theogenes Burmeister, 1847 -- invalid
	Thyce LeConte, 1856 -- valid
	Thyce angusticollis Casey, 1914 -- invalid
	Thyce angustula Casey, 1914 -- invalid
	Thyce aperta Casey, 1914 -- invalid
	Thyce aurata Van Dyke, 1943 -- invalid
	Thyce blaisdelli Casey, 1891 -- invalid
	Thyce brevitarsis Casey, 1914 -- invalid
	Thyce cinerea Van Dyke, 1943 -- invalid
	Thyce clypeata Van Dyke, 1943 -- invalid
	Thyce crinicollis Casey, 1914 -- invalid
	Thyce deserta Hardy, 1974 -- valid
	Thyce fossiger Casey, 1889 -- invalid
	Thyce longipalpis Casey, 1914 -- invalid
	Thyce nanella Casey, 1914 -- invalid
	Thyce nitidula Casey, 1914 -- invalid
	Thyce ochreata Casey, 1914 -- invalid
	Thyce pistoria Casey, 1895 -- invalid
	Thyce pulverea Casey, 1889 -- invalid
	Thyce rotundicauda Casey, 1914 -- invalid
	Thyce simplicipes Casey, 1914 -- invalid
	Thyce squamicollis LeConte, 1856 -- valid
	Thyce squamosa Casey, 1891 -- invalid
	Thyce vestita Casey, 1914 -- invalid
	Tomarus Erichson, 1847 -- valid
	Tomarus cuniculus (Fabricius, 1801) -- valid
	Tomarus gibbosus (De Geer, 1774) -- valid
	Tomarus laevicollis (Bates, 1888) -- valid
	Tomarus nasutus (Burmeister, 1847) -- valid
	Tomarus neglectus (LeConte, 1847) -- valid
	Tomarus relictus (Say, 1825) -- valid
	Tomarus sallaei (Bates, 1888) -- valid
	Tomarus selanderi (Cartwright, 1959) -- valid
	Tomarus subtropicus (Blatchley, 1922) -- valid
	Tostegoptera ventricosa LeConte, 1854 -- invalid
	Toxeutotaenius Kuwert, 1896 -- invalid
	Trichestes comans Burmeister, 1855 -- invalid
	Trichesthes Erichson, 1847 -- invalid
	Trichiini Fleming, 1821 -- valid
	Trichinus Kirby, 1827 -- invalid
	Trichiorhyssemus Clou�t, 1901 -- valid
	Trichiorhyssemus alternatus Hinton, 1938 -- valid
	Trichiorhyssemus cristatellus (Bates, 1887) -- valid
	Trichiorhyssemus riparius (Horn, 1871) -- valid
	Trichiotinus Casey, 1915 -- valid
	Trichiotinus affinis (Gory and Percheron, 1833) -- valid
	Trichiotinus assimilis (Kirby, 1837) -- valid
	Trichiotinus bibens (Fabricius, 1775) -- valid
	Trichiotinus intermedius Casey, 1915 -- invalid
	Trichiotinus lunulatus (Fabricius, 1775) -- valid
	Trichiotinus monticola Casey, 1915 -- invalid
	Trichiotinus piger (Fabricius, 1775) -- valid
	Trichiotinus reductus Casey, 1915 -- invalid
	Trichiotinus rufobrunneus (Casey, 1914) -- valid
	Trichiotinus texanus (Horn, 1876) -- valid
	Trichiotinus viridans (Kirby, 1837) -- valid
	Trichius bigsbii Kirby, 1827 -- invalid
	Trichius bistriga Newman, 1838 -- invalid
	Trichius carolinensis Casey, 1914 -- invalid
	Trichius drummond Gory and Percheron, 1833 -- invalid
	Trichius foveatus Kirby, 1837 -- invalid
	Trichius mutabilis Schaum, 1844 -- invalid
	Trichius obesulus Casey, 1914 -- invalid
	Trichius parvulus Casey, 1915 -- invalid
	Trichius rasilicaudus Casey, 1915 -- invalid
	Trichius rotundicollis Kirby, 1837 -- invalid
	Trichius rufiventris Casey, 1915 -- invalid
	Trichius rugosus Kirby, 1837 -- invalid
	Trichius semiviridis Casey, 1914 -- invalid
	Trichius squamiger Palisot de Beauvois, 1805 -- invalid
	Trichius variabilis Burmeister and Schaum, 1841 -- invalid
	Trichius variegatus Palisot de Beauvois, 1805 -- invalid
	Trichius venticosus Casey, 1915 -- invalid
	Trichius virens Gmelin, 1790 -- invalid
	Trichius viridulus Fabricius, 1775 -- invalid
	Trichoderma Nonfried, 1894 -- invalid
	Trichonotulus Bedel, 1911 -- valid
	Trichonotulus scrofa (Fabricius, 1787) -- valid
	Trichonotus Mulsant, 1842 -- invalid
	Trichonotus setiger Mulsant, 1842 -- invalid
	Trichopertha Reitter, 1903 -- invalid
	Trigonopeltastes Burmeister, 1840 -- valid
	Trigonopeltastes archimedes Schaum, 1841 -- valid
	Trigonopeltastes delta (Forster, 1771) -- valid
	Trigonopeltastes discrepans Howden, 1968 -- valid
	Trigonopeltastes floridana (Casey, 1909) -- invalid -- Scrub palmetto flower scarab
	Trigonopeltastes floridanus (Casey, 1909) -- valid
	Trigonopeltastes frontalis Bates, 1889 -- valid
	Trigonopeltastes sallaei sinaloensis Howden, 1968 -- valid
	Trigonopeltastes truncatus Howden, 1968 -- valid
	Trogidae MacLeay, 1819 -- valid
	Trox Fabricius, 1775 -- valid
	Trox aciculeatus Robinson, 1940 -- invalid
	Trox aequalis Say, 1832 -- valid
	Trox affinis Robinson, 1940 -- valid
	Trox alternans LeConte, 1854 -- invalid
	Trox alternatus Say, 1831 -- invalid
	Trox arenarius Fabricius, 1787 -- invalid
	Trox arenosus Gmelin, 1790 -- invalid
	Trox atrox LeConte, 1854 -- valid
	Trox barbosus von Laicharting, 1781 -- invalid
	Trox canaliculatus Say, 1824 -- invalid
	Trox capillaris Say, 1824 -- valid
	Trox confluentus Robinson, 1941 -- invalid
	Trox contractus Robinson, 1940 -- valid
	Trox crenatus Olivier, 1789 -- invalid
	Trox denticulatus Palisot de Beauvois, 1818 -- invalid
	Trox erinaceus LeConte, 1854 -- invalid
	Trox fascifer LeConte, 1854 -- valid
	Trox floridanus Howden and Vaurie, 1957 -- valid
	Trox foveicollis Harold, 1872 -- valid
	Trox frontera Vaurie, 1955 -- valid
	Trox gemmulatus Horn, 1874 -- valid
	Trox gibbus Olivier, 1789 -- invalid
	Trox hamatus Robinson, 1940 -- valid
	Trox hispidus von Paykull, 1798 -- invalid
	Trox howelli Howden and Vaurie, 1957 -- invalid -- Caracara commensal scarab
	Trox laticollis LeConte, 1854 -- valid
	Trox lecontei Harold, 1872 -- invalid
	Trox manilensis Schultze, 1915 -- invalid
	Trox monticola Robinson, 1940 -- invalid
	Trox nigrescens Robinson, 1941 -- invalid
	Trox niponensis Lewis, 1895 -- invalid
	Trox nobilis Wollaston, 1867 -- invalid
	Trox oligonus Loomis, 1922 -- invalid
	Trox ovatus Palisot de Beauvois, 1818 -- invalid
	Trox platycyphus Loomis, 1922 -- invalid
	Trox plicatus Robinson, 1940 -- valid
	Trox porcatus Say, 1825 -- invalid
	Trox robinsoni Vaurie, 1955 -- valid
	Trox scaber (Linnaeus, 1767) -- valid
	Trox serrulatus Palisot de Beauvois, 1818 -- invalid
	Trox sonorae LeConte, 1854 -- valid
	Trox sordidus LeConte, 1854 -- valid
	Trox spinulosus Robinson, 1940 -- valid
	Trox spinulosus dentibius Robinson, 1940 -- valid
	Trox spinulosus simi Robinson, 1940 -- valid
	Trox spinulosus spinulosus Robinson, 1940 -- valid
	Trox splendidus Say, 1835 -- invalid
	Trox striatus Melsheimer, 1845 -- valid
	Trox terrestris Say, 1825 -- valid
	Trox torressalai Baguena, 1959 -- invalid
	Trox tricolor Blackburn, 1904 -- invalid
	Trox triestinae Pittino, 1987 -- invalid
	Trox trisulcatus Curtis, 1844 -- invalid
	Trox tuberculatus Palisot de Beauvois, 1818 -- invalid
	Trox tuberculatus (De Geer, 1774) -- valid
	Trox unistriatus Palisot de Beauvois, 1818 -- valid
	Trox variolatus Melsheimer, 1845 -- valid
	Urleta Westwood, 1875 -- invalid
	Uroxys Westwood, 1842 -- valid
	Uroxys deavilai Delgado and Kohlmann, 2007 -- valid
	Valerius Kuwert, 1891 -- invalid
	Valgini Mulsant, 1842 -- valid
	Valgus Scriba, 1790 -- valid
	Valgus californicus Horn, 1870 -- valid
	Valgus canaliculatus (Olivier, 1789) -- valid
	Valgus hemipterus (Linnaeus, 1758) -- valid
	Valgus minutus Casey, 1909 -- invalid
	Valgus rubi B�guena, 1955 -- invalid
	Valgus rufosquamatus Dalla Torre, 1879 -- invalid
	Valgus serricollis Fitch, 1858 -- invalid
	Valgus seticollis (Palisot de Beauvois, 1805) -- valid
	Warwickia Smith and Evans, 2005 -- valid
	Warwickia pilosa (Sanderson, 1939) -- valid
	Xenochodaeus Paulsen, 2007 -- valid
	Xenochodaeus americanus (Westwood, 1852) -- valid
	Xenochodaeus luscinus (Howden, 1968) -- valid
	Xenochodaeus musculus (Say, 1835) -- valid
	Xenochodaeus planifrons (Schaeffer, 1906) -- valid
	Xenochodaeus simplex (LeConte, 1854) -- valid
	Xenochodaeus ulkei (Horn, 1876) -- valid
	Xenopopillia Kolbe, 1910 -- invalid
	Xeropsamobeus Saylor, 1937 -- valid
	Xeropsamobeus acerbus (Horn, 1887) -- valid
	Xeropsamobeus ambiguus (Fall, 1907) -- valid
	Xeropsamobeus arenicolus Gordon and Skelley, 2007 -- valid
	Xeropsamobeus asellus (Schmidt, 1907) -- valid
	Xeropsamobeus brighti Gordon and Skelley, 2007 -- valid
	Xeropsamobeus desertus (Van Dyke, 1918) -- valid
	Xeropsamobeus doyeni Gordon and Skelley, 2007 -- valid
	Xeropsamobeus mohavei Gordon and Skelley, 2007 -- valid
	Xeropsamobeus padrei Gordon and Skelley, 2007 -- valid
	Xeropsamobeus scabriceps (LeConte, 1878) -- valid
	Xyloryctes Hope, 1837 -- valid
	Xyloryctes ensifer Bates, 1888 -- valid
	Xyloryctes faunus Casey, 1895 -- invalid
	Xyloryctes hebes Casey, 1915 -- invalid
	Xyloryctes intermedia Sternberg, 1908 -- invalid
	Xyloryctes jamaicensis (Drury, 1773) -- valid
	Xyloryctes lacustris Casey, 1915 -- invalid
	Xyloryctes laevipennis Sternberg, 1908 -- invalid
	Xyloryctes obsolescens Casey, 1915 -- invalid
	Xyloryctes perpendicularis Sternberg, 1908 -- invalid
	Xyloryctes telephus Burmeister, 1847 -- valid
	Xyloryctes tenuicornutus Casey, 1915 -- invalid
	Xyloryctes thestalus Bates, 1888 -- valid
	Xyloryctes thestalus borealis Endr�di, 1975 -- valid
	Xyloryctes thestalus thestalus Bates, 1888 -- valid
	Zosterothrix Zang, 1905 -- invalid

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