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Heriades  Spinola, 1808
Taxonomic Serial No.: 154375

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 Taxonomy and Nomenclature
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Common Name(s):    
  Taxonomic Status:    
  Current Standing: valid  
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2021   

 Taxonomic Hierarchy
 KingdomAnimalia  – Animal, animaux, animals  
    SubkingdomBilateria  – triploblasts  
             PhylumArthropoda  – Artrópode, arthropodes, arthropods  
                SubphylumHexapoda  – hexapods  
                   ClassInsecta  – insects, hexapoda, inseto, insectes  
                      SubclassPterygota  – insects ailés, winged insects  
                         InfraclassNeoptera  – modern, wing-folding insects  
                               OrderHymenoptera  – abelha, formiga, vespa, ants, bees, wasps  
                                  SuborderApocrita  – abeilles, fourmis, guępes véritables, narrow-waisted hymenopterans, ants, bees, true wasps  
                                        SuperfamilyApoidea  – bees, sphecoid wasps, apoid wasps  
                                           FamilyMegachilidae  – leafcutting bees  
                                                    GenusHeriades Spinola, 1808  
    Direct Children:  
                                                       Species Heriades abessinica Friese, 1915  
                                                       Species Heriades albiscopana Strand, 1912  
                                                       Species Heriades aldabrana Cockerell, 1912  
                                                       Species Heriades alfkeni Benoist, 1938  
                                                       Species Heriades ambigua Cockerell, 1946  
                                                       Species Heriades ammodendri Popov, 1960  
                                                       Species Heriades angusticeps Cockerell, 1937  
                                                       Species Heriades angustula Cockerell, 1937  
                                                       Species Heriades apricula Griswold, 1998  
                                                       Species Heriades arcuatella Cockerell, 1937  
                                                       Species Heriades arnoldi Friese, 1922  
                                                       Species Heriades auroscopata (Gupta, Simlote and Sharma, 1993)  
                                                       Species Heriades bequaerti Cockerell, 1931  
                                                       Species Heriades bidentata (Gupta and Simlote, 1993)  
                                                       Species Heriades binodosa Cockerell, 1936  
                                                       Species Heriades bouyssoui Vachal, 1903  
                                                       Species Heriades brage Strand, 1912  
                                                       Species Heriades bruneri Titus, 1904  
                                                       Species Heriades burgeoni Benoist, 1931  
                                                       Species Heriades canaliculata Benoist, 1931  
                                                       Species Heriades cancava Wu, 1982  
                                                       Species Heriades capicola Strand, 1912  
                                                       Species Heriades carinata Cresson, 1864  
                                                       Species Heriades chariensis Benoist, 1931  
                                                       Species Heriades chrysogaster (Cameron, 1897)  
                                                       Species Heriades chubbi Cockerell, 1916  
                                                       Species Heriades cingulata Benoist, 1931  
                                                       Species Heriades civica Cockerell, 1937  
                                                       Species Heriades clavicornis Morawitz, 1875  
                                                       Species Heriades crenulata Nylander, 1856  
                                                       Species Heriades cressoni Michener, 1938  
                                                       Species Heriades crucifera Cockerell, 1897  
                                                       Species Heriades currani Michener, 1943  
                                                       Species Heriades curviventris Strand, 1912  
                                                       Species Heriades diminuta Cockerell, 1937  
                                                       Species Heriades discrepans Benoist, 1938  
                                                       Species Heriades edentatus (Friese, 1922)  
                                                       Species Heriades ellenbergeri Benoist, 1931  
                                                       Species Heriades erecta Gupta, 1987  
                                                       Species Heriades erythrosoma (Friese, 1922)  
                                                       Species Heriades eximia Friese, 1904  
                                                       Species Heriades fasciata (Gupta, Sharma and Simlote, 1993)  
                                                       Species Heriades fertoni Benoist, 1938  
                                                       Species Heriades filicornis Friese, 1915  
                                                       Species Heriades floccifera Brauns, 1929  
                                                       Species Heriades freygessneri Schletterer, 1889  
                                                       Species Heriades frontosa Schletterer, 1889  
                                                       Species Heriades fuelleborni Friese, 1922  
                                                       Species Heriades fujiyamai Yasumatsu and Hirashima, 1952  
                                                       Species Heriades fulvescens Cockerell, 1920  
                                                       Species Heriades fulvohispida Yasumatsu and Hirashima, 1952  
                                                       Species Heriades gibbosa (Friese, 1909)  
                                                       Species Heriades glomerans Schletterer, 1889  
                                                       Species Heriades gracilior Cockerell, 1897  
                                                       Species Heriades haryanensis (Gupta, Sharma and Simlote, 1993)  
                                                       Species Heriades hercules Strand, 1912  
                                                       Species Heriades hexadenticulata (Gupta, Sharma and Simlote, 1993)  
                                                       Species Heriades himalayensis Gupta, 1993  
                                                       Species Heriades hissarica Popov, 1955  
                                                       Species Heriades impressa Schletterer, 1889  
                                                       Species Heriades indica (Gupta, Sharma and Simlote, 1993)  
                                                       Species Heriades ingogoensis Cockerell, 1946  
                                                       Species Heriades labiata Pérez, 1895  
                                                       Species Heriades langenburgica Strand, 1911  
                                                       Species Heriades laosella Cockerell, 1929  
                                                       Species Heriades latipes Popov, 1960  
                                                       Species Heriades leavitti Crawford, 1913  
                                                       Species Heriades liberica Cockerell, 1931  
                                                       Species Heriades lobata (Gupta and Sharma, 1993)  
                                                       Species Heriades longicornis Friese, 1915  
                                                       Species Heriades longicornis Popov, 1960  
                                                       Species Heriades longispinis Cockerell, 1944  
                                                       Species Heriades macrognata (Vachal, 1909)  
                                                       Species Heriades mamillifera Brauns, 1929  
                                                       Species Heriades mandibularis Friese, 1922  
                                                       Species Heriades micheneri Timberlake, 1947  
                                                       Species Heriades micheneriana Engel, 2015  
                                                       Species Heriades micropthalma Michener, 1954  
                                                       Species Heriades microsticta Cockerell, 1936  
                                                       Species Heriades mundula Cockerell, 1920  
                                                       Species Heriades nasifera Cockerell, 1946  
                                                       Species Heriades nigroa (Gupta and Sharma, 1993)  
                                                       Species Heriades nitescens Cockerell, 1931  
                                                       Species Heriades nodulosa Cockerell, 1937  
                                                       Species Heriades nursei (Gupta and Sharma, 1993)  
                                                       Species Heriades occidentalis Michener, 1938  
                                                       Species Heriades orientalis Gupta, 1987  
                                                       Species Heriades ornaticornis Cockerell, 1936  
                                                       Species Heriades otavica Eardley and Urban, 2006  
                                                       Species Heriades othonis Friese, 1914  
                                                       Species Heriades pachyacantha Cockerell, 1940  
                                                       Species Heriades paganensis Yasumatsu, 1942  
                                                       Species Heriades parvula Bingham, 1897  
                                                       Species Heriades patella (Nurse, 1902)  
                                                       Species Heriades perminuta Cockerell, 1935  
                                                       Species Heriades phthisica Gerstäcker, 1858  
                                                       Species Heriades plumosa Krombein, 1950  
                                                       Species Heriades pogonura Benoist, 1931  
                                                       Species Heriades pretorii Cockerell, 1946  
                                                       Species Heriades prionsa (Cameron, 1905)  
                                                       Species Heriades psiadiae Pauly and Griswold, 2001  
                                                       Species Heriades punctulata Cockerell, 1917  
                                                       Species Heriades punctulifera Schletterer, 1889  
                                                       Species Heriades rowlandi (Cockerell, 1947)  
                                                       Species Heriades rubicola Pérez, 1890  
                                                       Species Heriades rufifrons Cockerell, 1932  
                                                       Species Heriades rufoflagellata Gupta and Simlote, 1993  
                                                       Species Heriades rufomarginata (Gupta, Sharma and Simlote, 1993)  
                                                       Species Heriades sakishimana Yasumatsu and Hirashima, 1965  
                                                       Species Heriades sauteri Cockerell, 1911  
                                                       Species Heriades schwarzi Griswold, 1998  
                                                       Species Heriades scutellatus Friese, 1922  
                                                       Species Heriades seyrigi Pauly and Griswold, 2001  
                                                       Species Heriades shestakovi Popov, 1960  
                                                       Species Heriades solanensis (Gupta, Simlote and Sharma, 1993)  
                                                       Species Heriades spiniscutis (Cameron, 1905)  
                                                       Species Heriades strictifrons Cockerell, 1946  
                                                       Species Heriades subfrontosa Cockerell, 1939  
                                                       Species Heriades sulcatifera Cockerell, 1937  
                                                       Species Heriades sulcatifrons Cockerell, 1936  
                                                       Species Heriades sulcatula Cockerell, 1931  
                                                       Species Heriades tayrona Gonzalez and Griswold, 2011  
                                                       Species Heriades tenuis Nurse, 1904  
                                                       Species Heriades testaceicornis (Cameron, 1908)  
                                                       Species Heriades texana Michener, 1938  
                                                       Species Heriades timberlakei Michener, 1938  
                                                       Species Heriades trigibbifera Brauns, 1929  
                                                       Species Heriades truncorum (Linnaeus, 1758)  
                                                       Species Heriades turcomanica Popov, 1955  
                                                       Species Heriades usakensis Cockerell, 1937  
                                                       Species Heriades ustulata Benoist, 1931  
                                                       Species Heriades variolosa (Cresson, 1872)  
                                                       Species Heriades victoriae Friese, 1922  
                                                       Species Heriades wellmani Cockerell, 1908  
                                                       Species Heriades wilmattae Cockerell, 1931  
                                                       Species Heriades xanthogaster Cockerell, 1932  
                                                       Species Heriades yunnanensis Wu, 1992  

  Expert: John S. Ascher, PhD  
  Notes: Assistant Professor, Department of Biological Sciences, National University of Singapore   
  Reference for: Heriades    
  Other Source(s):    
  Source: Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical..., website (version 03July2012)  
  Acquired: 2016   
  Notes: Moure, J. S., Urban, D., and G. A. R. Melo (Orgs). Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region. Available online at   
  Reference for: Heriades   
  Source: Discover Life bee species guide and world checklist - Draft-54, website (version Draft-54)  
  Acquired: 2020   
  Notes: Ascher, J. S. and J. Pickering. 1 February, 2020. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila).   
  Reference for: Heriades   
  Source: Hymenoptera Name Server version 0.03 4.x.2002, website (version 0.03)  
  Acquired: 2004   
  Reference for: Heriades   
  Source: NODC Taxonomic Code, database (version 8.0)  
  Acquired: 1996   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.  
  Publication Date: 2000   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World   
  Page(s): xiv + 913   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-6133-0   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.  
  Publication Date: 2007   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World, Second Edition   
  Page(s): xvi + 953   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-8573-6   
  Reference for: Heriades   
  Author(s)/Editor(s): Spinola, M.  
  Publication Date: 1808   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Insectorum Liguriae Species Novae aut Rariores, quas in agro Ligustico nuper detexit, descripsit, et iconibus illustravit Maximilianus Spinola, adjecto Catalogo spiecierum auctoribus jam enumeratarum, quae in eadam regione occurrunt, vol. 2   
  Page(s): ii + 262   
  Publisher: Gravier   
  Publication Place: Genuae (Genoa), Italy   
  Notes: Available online via Biodiversity Heritage Library at   
  Reference for: original description   
  Author(s)/Editor(s): Ungricht, Stefan, Andreas Müller, and Silvia Dorn  
  Publication Date: 2008   
  Article/Chapter Title: A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)   
  Journal/Book Name, Vol. No.: Zootaxa, issue 1865   
  Page(s): 253   
  Publication Place:    
  ISBN/ISSN: 9781869772550, 1175-5326  
  Notes: Online edition ISBN is 978-1-86977-256-7 and ISSN is 1175-5334:   
  Reference for: Heriades   

 Geographic Information
  Geographic Division:    



  Comment: The gender of Heriades Spinola 1808 is feminine. Spinola (1808: 7) proposed Heriades, and included several species without giving explicit combinations in the new genus. However, he did (p.9) present two new species names, as 'Heriades pusilla, [H.] sinuata, mihi,' which clearly indicates his use of the genus name as feminine. An etymological commentary by Doug Yanega (Yahoo BeeMonitoring Listserv, 5 October, 2015) is here noted, for reference: 'Dalla Torre and Friese were absolutely wrong; there is no form of the noun for wool - either "eiros" or "erion" - which would be spelled as "Eriades". The term "eriodes" would mean "woolly". If you look at Spinola, his entire paper was written in Latin, and he uses the forms "Heriadibus" and "Heriadum", which indicate that the word "Heriades" is a 3rd-declension plural noun, so the singular form would be "Heriad" - either a son of Eris (which would be masculine), or a mountain nymph (which would be feminine). He included two feminine species, so we know which of the two he intended'  



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