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Heriades  Spinola, 1808
Taxonomic Serial No.: 154375

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 Taxonomy and Nomenclature
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Common Name(s):    
  Taxonomic Status:    
  Current Standing: valid  
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2021   

 Taxonomic Hierarchy
 KingdomAnimalia  – Animal, animaux, animals  
             PhylumArthropoda  – Artrópode, arthropodes, arthropods  
                SubphylumHexapoda  – hexapods  
                   ClassInsecta  – insects, hexapoda, inseto, insectes  
                      SubclassPterygota  – insects ailés, winged insects  
                         InfraclassNeoptera  – modern, wing-folding insects  
                               OrderHymenoptera  – abelha, formiga, vespa, ants, bees, wasps  
                                  SuborderApocrita  – abeilles, fourmis, guępes véritables, narrow-waisted hymenopterans, ants, bees, true wasps  
                                        SuperfamilyApoidea  – bees, sphecoid wasps, apoid wasps  
                                           FamilyMegachilidae  – leafcutting bees  
                                                    GenusHeriades Spinola, 1808  
    Direct Children:  
                                                       SpeciesHeriades abessinica Friese, 1915  
                                                       SpeciesHeriades albiscopana Strand, 1912  
                                                       SpeciesHeriades aldabrana Cockerell, 1912  
                                                       SpeciesHeriades alfkeni Benoist, 1938  
                                                       SpeciesHeriades ambigua Cockerell, 1946  
                                                       SpeciesHeriades ammodendri Popov, 1960  
                                                       SpeciesHeriades angusticeps Cockerell, 1937  
                                                       SpeciesHeriades angustula Cockerell, 1937  
                                                       SpeciesHeriades apricula Griswold, 1998  
                                                       SpeciesHeriades arcuatella Cockerell, 1937  
                                                       SpeciesHeriades arnoldi Friese, 1922  
                                                       SpeciesHeriades auroscopata (Gupta, Simlote and Sharma, 1993)  
                                                       SpeciesHeriades bequaerti Cockerell, 1931  
                                                       SpeciesHeriades bidentata (Gupta and Simlote, 1993)  
                                                       SpeciesHeriades binodosa Cockerell, 1936  
                                                       SpeciesHeriades bouyssoui Vachal, 1903  
                                                       SpeciesHeriades brage Strand, 1912  
                                                       SpeciesHeriades bruneri Titus, 1904  
                                                       SpeciesHeriades burgeoni Benoist, 1931  
                                                       SpeciesHeriades canaliculata Benoist, 1931  
                                                       SpeciesHeriades cancava Wu, 1982  
                                                       SpeciesHeriades capicola Strand, 1912  
                                                       SpeciesHeriades carinata Cresson, 1864  
                                                       SpeciesHeriades chariensis Benoist, 1931  
                                                       SpeciesHeriades chrysogaster (Cameron, 1897)  
                                                       SpeciesHeriades chubbi Cockerell, 1916  
                                                       SpeciesHeriades cingulata Benoist, 1931  
                                                       SpeciesHeriades civica Cockerell, 1937  
                                                       SpeciesHeriades clavicornis Morawitz, 1875  
                                                       SpeciesHeriades crenulata Nylander, 1856  
                                                       SpeciesHeriades cressoni Michener, 1938  
                                                       SpeciesHeriades crucifera Cockerell, 1897  
                                                       SpeciesHeriades currani Michener, 1943  
                                                       SpeciesHeriades curviventris Strand, 1912  
                                                       SpeciesHeriades diminuta Cockerell, 1937  
                                                       SpeciesHeriades discrepans Benoist, 1938  
                                                       SpeciesHeriades edentatus (Friese, 1922)  
                                                       SpeciesHeriades ellenbergeri Benoist, 1931  
                                                       SpeciesHeriades erecta Gupta, 1987  
                                                       SpeciesHeriades erythrosoma (Friese, 1922)  
                                                       SpeciesHeriades eximia Friese, 1904  
                                                       SpeciesHeriades fasciata (Gupta, Sharma and Simlote, 1993)  
                                                       SpeciesHeriades fertoni Benoist, 1938  
                                                       SpeciesHeriades filicornis Friese, 1915  
                                                       SpeciesHeriades floccifera Brauns, 1929  
                                                       SpeciesHeriades freygessneri Schletterer, 1889  
                                                       SpeciesHeriades frontosa Schletterer, 1889  
                                                       SpeciesHeriades fuelleborni Friese, 1922  
                                                       SpeciesHeriades fujiyamai Yasumatsu and Hirashima, 1952  
                                                       SpeciesHeriades fulvescens Cockerell, 1920  
                                                       SpeciesHeriades fulvohispida Yasumatsu and Hirashima, 1952  
                                                       SpeciesHeriades gibbosa (Friese, 1909)  
                                                       SpeciesHeriades glomerans Schletterer, 1889  
                                                       SpeciesHeriades gracilior Cockerell, 1897  
                                                       SpeciesHeriades haryanensis (Gupta, Sharma and Simlote, 1993)  
                                                       SpeciesHeriades hercules Strand, 1912  
                                                       SpeciesHeriades hexadenticulata (Gupta, Sharma and Simlote, 1993)  
                                                       SpeciesHeriades himalayensis Gupta, 1993  
                                                       SpeciesHeriades hissarica Popov, 1955  
                                                       SpeciesHeriades impressa Schletterer, 1889  
                                                       SpeciesHeriades indica (Gupta, Sharma and Simlote, 1993)  
                                                       SpeciesHeriades ingogoensis Cockerell, 1946  
                                                       SpeciesHeriades labiata Pérez, 1895  
                                                       SpeciesHeriades langenburgica Strand, 1911  
                                                       SpeciesHeriades laosella Cockerell, 1929  
                                                       SpeciesHeriades latipes Popov, 1960  
                                                       SpeciesHeriades leavitti Crawford, 1913  
                                                       SpeciesHeriades liberica Cockerell, 1931  
                                                       SpeciesHeriades lobata (Gupta and Sharma, 1993)  
                                                       SpeciesHeriades longicornis Friese, 1915  
                                                       SpeciesHeriades longicornis Popov, 1960  
                                                       SpeciesHeriades longispinis Cockerell, 1944  
                                                       SpeciesHeriades macrognata (Vachal, 1909)  
                                                       SpeciesHeriades mamillifera Brauns, 1929  
                                                       SpeciesHeriades mandibularis Friese, 1922  
                                                       SpeciesHeriades micheneri Timberlake, 1947  
                                                       SpeciesHeriades micheneriana Engel, 2015  
                                                       SpeciesHeriades micropthalma Michener, 1954  
                                                       SpeciesHeriades microsticta Cockerell, 1936  
                                                       SpeciesHeriades mundula Cockerell, 1920  
                                                       SpeciesHeriades nasifera Cockerell, 1946  
                                                       SpeciesHeriades nigroa (Gupta and Sharma, 1993)  
                                                       SpeciesHeriades nitescens Cockerell, 1931  
                                                       SpeciesHeriades nodulosa Cockerell, 1937  
                                                       SpeciesHeriades nursei (Gupta and Sharma, 1993)  
                                                       SpeciesHeriades occidentalis Michener, 1938  
                                                       SpeciesHeriades orientalis Gupta, 1987  
                                                       SpeciesHeriades ornaticornis Cockerell, 1936  
                                                       SpeciesHeriades otavica Eardley and Urban, 2006  
                                                       SpeciesHeriades othonis Friese, 1914  
                                                       SpeciesHeriades pachyacantha Cockerell, 1940  
                                                       SpeciesHeriades paganensis Yasumatsu, 1942  
                                                       SpeciesHeriades parvula Bingham, 1897  
                                                       SpeciesHeriades patella (Nurse, 1902)  
                                                       SpeciesHeriades perminuta Cockerell, 1935  
                                                       SpeciesHeriades phthisica Gerstäcker, 1858  
                                                       SpeciesHeriades plumosa Krombein, 1950  
                                                       SpeciesHeriades pogonura Benoist, 1931  
                                                       SpeciesHeriades pretorii Cockerell, 1946  
                                                       SpeciesHeriades prionsa (Cameron, 1905)  
                                                       SpeciesHeriades psiadiae Pauly and Griswold, 2001  
                                                       SpeciesHeriades punctulata Cockerell, 1917  
                                                       SpeciesHeriades punctulifera Schletterer, 1889  
                                                       SpeciesHeriades rowlandi (Cockerell, 1947)  
                                                       SpeciesHeriades rubicola Pérez, 1890  
                                                       SpeciesHeriades rufifrons Cockerell, 1932  
                                                       SpeciesHeriades rufoflagellata Gupta and Simlote, 1993  
                                                       SpeciesHeriades rufomarginata (Gupta, Sharma and Simlote, 1993)  
                                                       SpeciesHeriades sakishimana Yasumatsu and Hirashima, 1965  
                                                       SpeciesHeriades sauteri Cockerell, 1911  
                                                       SpeciesHeriades schwarzi Griswold, 1998  
                                                       SpeciesHeriades scutellatus Friese, 1922  
                                                       SpeciesHeriades seyrigi Pauly and Griswold, 2001  
                                                       SpeciesHeriades shestakovi Popov, 1960  
                                                       SpeciesHeriades solanensis (Gupta, Simlote and Sharma, 1993)  
                                                       SpeciesHeriades spiniscutis (Cameron, 1905)  
                                                       SpeciesHeriades strictifrons Cockerell, 1946  
                                                       SpeciesHeriades subfrontosa Cockerell, 1939  
                                                       SpeciesHeriades sulcatifera Cockerell, 1937  
                                                       SpeciesHeriades sulcatifrons Cockerell, 1936  
                                                       SpeciesHeriades sulcatula Cockerell, 1931  
                                                       SpeciesHeriades tayrona Gonzalez and Griswold, 2011  
                                                       SpeciesHeriades tenuis Nurse, 1904  
                                                       SpeciesHeriades testaceicornis (Cameron, 1908)  
                                                       SpeciesHeriades texana Michener, 1938  
                                                       SpeciesHeriades timberlakei Michener, 1938  
                                                       SpeciesHeriades trigibbifera Brauns, 1929  
                                                       SpeciesHeriades truncorum (Linnaeus, 1758)  
                                                       SpeciesHeriades turcomanica Popov, 1955  
                                                       SpeciesHeriades usakensis Cockerell, 1937  
                                                       SpeciesHeriades ustulata Benoist, 1931  
                                                       SpeciesHeriades variolosa (Cresson, 1872)  
                                                       SpeciesHeriades victoriae Friese, 1922  
                                                       SpeciesHeriades wellmani Cockerell, 1908  
                                                       SpeciesHeriades wilmattae Cockerell, 1931  
                                                       SpeciesHeriades xanthogaster Cockerell, 1932  
                                                       SpeciesHeriades yunnanensis Wu, 1992  

  Expert: John S. Ascher, PhD   
  Notes: Assistant Professor, Department of Biological Sciences, National University of Singapore   
  Reference for: Heriades    
  Other Source(s):    
  Source: Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical..., website (version 03July2012)  
  Acquired: 2016   
  Notes: Moure, J. S., Urban, D., and G. A. R. Melo (Orgs). Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region. Available online at   
  Reference for: Heriades   
  Source: Discover Life bee species guide and world checklist - Draft-54, website (version Draft-54)  
  Acquired: 2020   
  Notes: Ascher, J. S. and J. Pickering. 1 February, 2020. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila).   
  Reference for: Heriades   
  Source: Hymenoptera Name Server version 0.03 4.x.2002, website (version 0.03)  
  Acquired: 2004   
  Reference for: Heriades   
  Source: NODC Taxonomic Code, database (version 8.0)  
  Acquired: 1996   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.   
  Publication Date: 2000   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World   
  Page(s): xiv + 913   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-6133-0   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.   
  Publication Date: 2007   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World, Second Edition   
  Page(s): xvi + 953   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-8573-6   
  Reference for: Heriades   
  Author(s)/Editor(s): Spinola, M.   
  Publication Date: 1808   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Insectorum Liguriae Species Novae aut Rariores, quas in agro Ligustico nuper detexit, descripsit, et iconibus illustravit Maximilianus Spinola, adjecto Catalogo spiecierum auctoribus jam enumeratarum, quae in eadam regione occurrunt, vol. 2   
  Page(s): ii + 262   
  Publisher: Gravier   
  Publication Place: Genuae (Genoa), Italy   
  Notes: Available online via Biodiversity Heritage Library at   
  Reference for: original description   
  Author(s)/Editor(s): Ungricht, Stefan, Andreas Müller, and Silvia Dorn   
  Publication Date: 2008   
  Article/Chapter Title: A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)   
  Journal/Book Name, Vol. No.: Zootaxa, issue 1865   
  Page(s): 253   
  Publication Place:    
  ISBN/ISSN: 9781869772550, 1175-5326  
  Notes: Online edition ISBN is 978-1-86977-256-7 and ISSN is 1175-5334:   
  Reference for: Heriades   

 Geographic Information
  Geographic Division:    



  Comment: The gender of Heriades Spinola 1808 is feminine. Spinola (1808: 7) proposed Heriades, and included several species without giving explicit combinations in the new genus. However, he did (p.9) present two new species names, as 'Heriades pusilla, [H.] sinuata, mihi,' which clearly indicates his use of the genus name as feminine. An etymological commentary by Doug Yanega (Yahoo BeeMonitoring Listserv, 5 October, 2015) is here noted, for reference: 'Dalla Torre and Friese were absolutely wrong; there is no form of the noun for wool - either "eiros" or "erion" - which would be spelled as "Eriades". The term "eriodes" would mean "woolly". If you look at Spinola, his entire paper was written in Latin, and he uses the forms "Heriadibus" and "Heriadum", which indicate that the word "Heriades" is a 3rd-declension plural noun, so the singular form would be "Heriad" - either a son of Eris (which would be masculine), or a mountain nymph (which would be feminine). He included two feminine species, so we know which of the two he intended'  



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