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Pteropus  Brisson, 1762
Taxonomic Serial No.: 180032

(Download Help) Pteropus TSN 180032

 Taxonomy and Nomenclature
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Common Name(s): flying foxes [English]
  Taxonomic Status:    
  Current Standing: valid  
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2020   

 Taxonomic Hierarchy
 KingdomAnimalia  – Animal, animaux, animals  
          PhylumChordata  – cordés, cordado, chordates  
             SubphylumVertebrata  – vertebrado, vertébrés, vertebrates  
                      ClassMammalia Linnaeus, 1758 – mammifères, mamífero, mammals  
                         SubclassTheria Parker and Haswell, 1897  
                            InfraclassEutheria Gill, 1872  
                               OrderChiroptera Blumenbach, 1779 – morcego, quiróptero, bats  
                                  SuborderYinpterochiroptera Springer, Teeling, Madsen, Stanhope and Jong, 2001  
                                     FamilyPteropodidae Gray, 1821 – flying foxes, Old World fruit bats, megabats, non-echolocating bats  
                                        SubfamilyPteropodinae Gray, 1821 – Old World fruit bats  
                                           TribePteropodini Gray, 1821  
                                              GenusPteropus Brisson, 1762 – flying foxes  
    Direct Children:  
                                                 Species Pteropus admiralitatum Thomas, 1894 – Admiralty Flying Fox 
                                                 Species Pteropus aldabrensis True, 1893 – Aldabra Flying Fox 
                                                 Species Pteropus alecto Temminck, 1837 – Black Flying Fox 
                                                 Species Pteropus allenorum Helgen, Helgen and Wilson, 2009 – Small Samoan Flying Fox 
                                                 Species Pteropus anetianus (Gray, 1870) – Vanauatu flying fox, Vanuatu Flying Fox 
                                                 Species Pteropus aruensis Peters, 1867 – Aru Flying Fox 
                                                 Species Pteropus brunneus Dobson, 1878 – Dusky Flying Fox 
                                                 Species Pteropus caniceps Gray, 1870 – North Moluccan Flying Fox 
                                                 Species Pteropus capistratus Peters, 1876 – Bismark Masked Flying Fox, New Britain Masked Flying Fox 
                                                 Species Pteropus chrysoproctus Temminck, 1837 – Moluccan Flying Fox, South Moluccan Flying Fox 
                                                 Species Pteropus cognatus K. Andersen, 1908 – Makira Flying Fox 
                                                 Species Pteropus conspicillatus Gould, 1850 – Spectacled Flying Fox 
                                                 Species Pteropus coxi Helgen, Helgen and Wilson, 2009 – Large Samoan Flying Fox 
                                                 Species Pteropus dasymallus Temminck, 1825 – Ryukyu Flying Fox 
                                                 Species Pteropus ennisae Flannery and White, 1991 – New Ireland Masked Flying Fox 
                                                 Species Pteropus faunulus Miller, 1902 – Nicobar Flying Fox 
                                                 Species Pteropus fundatus Felten and Kock, 1972 – Banks Flying Fox 
                                                 Species Pteropus gilliardorum Van Deusen, 1969 – Gilliard's Flying Fox 
                                                 Species Pteropus griseus (E. Geoffroy, 1810) – Gray Flying Fox, Wallacean Gray Flying Fox 
                                                 Species Pteropus howensis Troughton, 1931 – Ontong Java Flying Fox 
                                                 Species Pteropus hypomelanus Temminck, 1853 – Variable Flying Fox, Island Flying Fox 
                                                 Species Pteropus keyensis Peters, 1867 – Kei Flying Fox, Kai Flying Fox 
                                                 Species Pteropus livingstonii Gray, 1866 – Comoro Black Flying Fox, Comoro Flying Fox, Livingstone's Flying Fox 
                                                 Species Pteropus lombocensis Dobson, 1878 – Lombok Flying Fox 
                                                 Species Pteropus lylei K. Andersen, 1908 – Lyle's Flying Fox 
                                                 Species Pteropus macrotis Peters, 1867 – Big-eared Flying Fox 
                                                 Species Pteropus mahaganus Sanborn, 1931 – Sanborn's Flying Fox 
                                                 Species Pteropus mariannus Desmarest, 1822 – Mariana Flying Fox, Mariana Fruit Bat, Marianas Flying Fox 
                                                 Species Pteropus medius Temminck, 1825 – Indian Flying Fox 
                                                 Species Pteropus melanopogon Peters, 1867 – Black-bearded Flying Fox 
                                                 Species Pteropus melanotus Blyth, 1863 – Black-eared Flying Fox 
                                                 Species Pteropus molossinus Temminck, 1853 – Ponape Flying Fox, Caroline Flying Fox, Pohnpei Flying Fox 
                                                 Species Pteropus neohibernicus Peters, 1876 – Great Flying Fox 
                                                 Species Pteropus niger (Kerr, 1792) – Greater Mascarene Flying Fox 
                                                 Species Pteropus nitendiensis Sanborn, 1930 – Temotu Flying Fox 
                                                 Species Pteropus ocularis Peters, 1867 – Ceram Flying Fox, Seram Flying Fox 
                                                 Species Pteropus ornatus Gray, 1870 – Ornate Flying Fox 
                                                 Species Pteropus pelagicus Kittlitz in Lutké, 1836 – Chuuk Flying Fox, Truk Flying Fox, Ruck Flying Fox 
                                                 Species Pteropus pelewensis K. Andersen, 1908 – Pelew Flying Fox, Palau Flying Fox 
                                                 Species Pteropus personatus Temminck, 1825 – Masked Flying Fox, Moluccan Masked Flying Fox 
                                                 Species Pteropus pilosus K. Andersen, 1908 – large Palau flying fox, Palau flying fox, Large Pelew Flying Fox 
                                                 Species Pteropus pohlei Stein, 1933 – Geelvink Bay Flying Fox 
                                                 Species Pteropus poliocephalus Temminck, 1825 – Gray-headed Flying Fox 
                                                 Species Pteropus pselaphon Lay, 1829 – Bonin Flying Fox 
                                                 Species Pteropus pumilus Miller, 1910 – Little Golden-mantled Flying Fox 
                                                 Species Pteropus rayneri Gray, 1870 – Solomons Flying Fox 
                                                 Species Pteropus rennelli Troughton, 1929 – Rennell Flying Fox 
                                                 Species Pteropus rodricensis Dobson, 1878 – Rodriguez Flying Fox, Rodrigues Fruit Bat, Rodrigues Island Flying Fox, Rodrigues Flying Fox 
                                                 Species Pteropus rufus E. Geoffroy, 1803 – Madagascan Flying Fox, Malagasy Flying Fox 
                                                 Species Pteropus samoensis Peale, 1848 – Samoa Flying Fox, Samoan Flying Fox, Common Samoan Flying Fox 
                                                 Species Pteropus scapulatus Peters, 1862 – Little Red Flying Fox 
                                                 Species Pteropus seychellensis A. Milne-Edwards, 1878 – Seychelles Flying Fox 
                                                 Species Pteropus speciosus K. Andersen, 1908 – Philippine Gray Flying Fox 
                                                 Species Pteropus subniger (Kerr, 1792) – Dark Flying Fox 
                                                 Species Pteropus temminckii Peters, 1867 – Temminck's Flying Fox 
                                                 Species Pteropus tokudae Tate, 1934 – Tokuda's Flying Fox, Guam Flying Fox, Little Mariana Fruit Bat 
                                                 Species Pteropus tonganus Quoy and Gaimard, 1830 – Insular Flying Fox, Pacific Flying Fox 
                                                 Species Pteropus tuberculatus Peters, 1869 – Vanikoro Flying Fox 
                                                 Species Pteropus ualanus Peters, 1883 – Kosrae Flying Fox 
                                                 Species Pteropus vampyrus (Linnaeus, 1758) – Large Flying Fox 
                                                 Species Pteropus vetula Jouan, 1863 – New Caledonia Flying Fox 
                                                 Species Pteropus voeltzkowi Matschie, 1909 – Pemba Flying Fox 
                                                 Species Pteropus woodfordi Thomas, 1888 – Dwarf Flying Fox 

  Expert: Alfred L. Gardner  
  Notes: Curator of North American mammals and Chief of Mammal Section, National Biological Service, Smithsonian Institution, National Museum of Natural History, Washington, DC, USA   
  Reference for: Pteropus    
  Expert: Don E. Wilson  
  Notes: Smithsonian Institution, MRC 108 Washington DC 20560-0108   
  Reference for: Pteropus    
  Other Source(s):    
  Source: Checklist of CITES species: Index of CITES species, document (version 2013)  
  Acquired: 2013   
  Notes: UNEP-WCMC (Comps.) 2013. Checklist of CITES species. CITES Secretariat, Geneva, Switzerland, and UNEP-WCMC, Cambridge, United Kingdom. Available online at   
  Reference for: Pteropus   
  Source: NODC Taxonomic Code, database (version 8.0)  
  Acquired: 1996   
  Reference for: Pteropus   
  Author(s)/Editor(s): Banks, R. C., R. W. McDiarmid, and A. L. Gardner  
  Publication Date: 1987   
  Article/Chapter Title: Checklist of Vertebrates of the United States, the U.S. Territories, and Canada   
  Journal/Book Name, Vol. No.: Resource Publication, no. 166   
  Page(s): 79   
  Publisher: United States Department of the Interior Fish and Wildlife Service   
  Publication Place: Washington, D.C., USA   
  Reference for: Pteropus   
  Author(s)/Editor(s): Simmons, Nancy B. / Wilson, Don E., and DeeAnn M. Reeder, eds.  
  Publication Date: 2005   
  Article/Chapter Title: Order Chiroptera   
  Journal/Book Name, Vol. No.: Mammal Species of the World: A Taxonomic and Geographic Reference, 3rd ed., vols. 1 & 2   
  Page(s): 312-529   
  Publisher: Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-8221-4   
  Notes: Corrections were made to text at 3rd printing   
  Reference for: Pteropus   
  Author(s)/Editor(s): Wilson, D. E., and R. A. Mittermeier, eds.  
  Publication Date: 2019   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Handbook of the Mammals of the World. Vol. 9: Bats   
  Page(s): 1008   
  Publisher: Lynx Edicions   
  Publication Place: Barcelona, Spain   
  ISBN/ISSN: 978-84-16728190   
  Reference for: Pteropus   
  Author(s)/Editor(s): Wilson, Don E., and DeeAnn M. Reeder, eds.  
  Publication Date: 1993   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Mammal Species of the World: A Taxonomic and Geographic Reference, 2nd ed., 3rd printing   
  Page(s): xviii + 1207   
  Publisher: Smithsonian Institution Press   
  Publication Place: Washington, DC, USA   
  ISBN/ISSN: 1-56098-217-9   
  Notes: Corrections were made to text at 3rd printing   
  Reference for: Pteropus   

 Geographic Information
  Geographic Division:    



  Comment: Comments: The International Commission on Zoological Nomenclature ruled in favor of rejecting Brisson (1762), but conserved several of the generic names including Pteropus (Opinion 1894, ICZN 1998). Formerly included arquatus and leucotis which were transferred to Acerodon by Musser et al. (1982a). Species groups follow Koopman (1994)  
    CITES: Appendix II except Pteropus brunneus and the species included in Appendix I, per 2013 Checklist of CITES species: Index of CITES species published with amendments to the Appendices and changes in standard scientific nomenclature following the 16th meeting of the Conference of the Parties (Bangkok, 2013). For current status see  



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Disclaimer: ITIS taxonomy is based on the latest scientific consensus available, and is provided as a general reference source for interested parties. However, it is not a legal authority for statutory or regulatory purposes. While every effort has been made to provide the most reliable and up-to-date information available, ultimate legal requirements with respect to species are contained in provisions of treaties to which the United States is a party, wildlife statutes, regulations, and any applicable notices that have been published in the Federal Register. For further information on U.S. legal requirements with respect to protected taxa, please contact the U.S. Fish and Wildlife Service.

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