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Pteropus  Brisson, 1762
Taxonomic Serial No.: 180032

(Download Help) Pteropus TSN 180032

 Taxonomy and Nomenclature
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Common Name(s): flying foxes [English]
  Taxonomic Status:    
  Current Standing: valid  
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2020   

 Taxonomic Hierarchy
 KingdomAnimalia  – Animal, animaux, animals  
          PhylumChordata  – cordés, cordado, chordates  
             SubphylumVertebrata  – vertebrado, vertébrés, vertebrates  
                      ClassMammalia Linnaeus, 1758 – mammifères, mamífero, mammals  
                         SubclassTheria Parker and Haswell, 1897  
                            InfraclassEutheria Gill, 1872  
                               OrderChiroptera Blumenbach, 1779 – morcego, quiróptero, bats  
                                  SuborderYinpterochiroptera Springer, Teeling, Madsen, Stanhope and Jong, 2001  
                                     FamilyPteropodidae Gray, 1821 – flying foxes, Old World fruit bats, megabats, non-echolocating bats  
                                        SubfamilyPteropodinae Gray, 1821 – Old World fruit bats  
                                           TribePteropodini Gray, 1821  
                                              GenusPteropus Brisson, 1762 – flying foxes  
    Direct Children:  
                                                 SpeciesPteropus admiralitatum Thomas, 1894 – Admiralty Flying Fox 
                                                 SpeciesPteropus aldabrensis True, 1893 – Aldabra Flying Fox 
                                                 SpeciesPteropus alecto Temminck, 1837 – Black Flying Fox 
                                                 SpeciesPteropus allenorum Helgen, Helgen and Wilson, 2009 – Small Samoan Flying Fox 
                                                 SpeciesPteropus anetianus (Gray, 1870) – Vanauatu flying fox, Vanuatu Flying Fox 
                                                 SpeciesPteropus aruensis Peters, 1867 – Aru Flying Fox 
                                                 SpeciesPteropus brunneus Dobson, 1878 – Dusky Flying Fox 
                                                 SpeciesPteropus caniceps Gray, 1870 – North Moluccan Flying Fox 
                                                 SpeciesPteropus capistratus Peters, 1876 – Bismark Masked Flying Fox, New Britain Masked Flying Fox 
                                                 SpeciesPteropus chrysoproctus Temminck, 1837 – Moluccan Flying Fox, South Moluccan Flying Fox 
                                                 SpeciesPteropus cognatus K. Andersen, 1908 – Makira Flying Fox 
                                                 SpeciesPteropus conspicillatus Gould, 1850 – Spectacled Flying Fox 
                                                 SpeciesPteropus coxi Helgen, Helgen and Wilson, 2009 – Large Samoan Flying Fox 
                                                 SpeciesPteropus dasymallus Temminck, 1825 – Ryukyu Flying Fox 
                                                 SpeciesPteropus ennisae Flannery and White, 1991 – New Ireland Masked Flying Fox 
                                                 SpeciesPteropus faunulus Miller, 1902 – Nicobar Flying Fox 
                                                 SpeciesPteropus fundatus Felten and Kock, 1972 – Banks Flying Fox 
                                                 SpeciesPteropus gilliardorum Van Deusen, 1969 – Gilliard's Flying Fox 
                                                 SpeciesPteropus griseus (E. Geoffroy, 1810) – Gray Flying Fox, Wallacean Gray Flying Fox 
                                                 SpeciesPteropus howensis Troughton, 1931 – Ontong Java Flying Fox 
                                                 SpeciesPteropus hypomelanus Temminck, 1853 – Variable Flying Fox, Island Flying Fox 
                                                 SpeciesPteropus keyensis Peters, 1867 – Kei Flying Fox, Kai Flying Fox 
                                                 SpeciesPteropus livingstonii Gray, 1866 – Comoro Black Flying Fox, Comoro Flying Fox, Livingstone's Flying Fox 
                                                 SpeciesPteropus lombocensis Dobson, 1878 – Lombok Flying Fox 
                                                 SpeciesPteropus lylei K. Andersen, 1908 – Lyle's Flying Fox 
                                                 SpeciesPteropus macrotis Peters, 1867 – Big-eared Flying Fox 
                                                 SpeciesPteropus mahaganus Sanborn, 1931 – Sanborn's Flying Fox 
                                                 SpeciesPteropus mariannus Desmarest, 1822 – Mariana Flying Fox, Mariana Fruit Bat, Marianas Flying Fox 
                                                 SpeciesPteropus medius Temminck, 1825 – Indian Flying Fox 
                                                 SpeciesPteropus melanopogon Peters, 1867 – Black-bearded Flying Fox 
                                                 SpeciesPteropus melanotus Blyth, 1863 – Black-eared Flying Fox 
                                                 SpeciesPteropus molossinus Temminck, 1853 – Ponape Flying Fox, Caroline Flying Fox, Pohnpei Flying Fox 
                                                 SpeciesPteropus neohibernicus Peters, 1876 – Great Flying Fox 
                                                 SpeciesPteropus niger (Kerr, 1792) – Greater Mascarene Flying Fox 
                                                 SpeciesPteropus nitendiensis Sanborn, 1930 – Temotu Flying Fox 
                                                 SpeciesPteropus ocularis Peters, 1867 – Ceram Flying Fox, Seram Flying Fox 
                                                 SpeciesPteropus ornatus Gray, 1870 – Ornate Flying Fox 
                                                 SpeciesPteropus pelagicus Kittlitz in Lutké, 1836 – Chuuk Flying Fox, Truk Flying Fox, Ruck Flying Fox 
                                                 SpeciesPteropus pelewensis K. Andersen, 1908 – Pelew Flying Fox, Palau Flying Fox 
                                                 SpeciesPteropus personatus Temminck, 1825 – Masked Flying Fox, Moluccan Masked Flying Fox 
                                                 SpeciesPteropus pilosus K. Andersen, 1908 – large Palau flying fox, Palau flying fox, Large Pelew Flying Fox 
                                                 SpeciesPteropus pohlei Stein, 1933 – Geelvink Bay Flying Fox 
                                                 SpeciesPteropus poliocephalus Temminck, 1825 – Gray-headed Flying Fox 
                                                 SpeciesPteropus pselaphon Lay, 1829 – Bonin Flying Fox 
                                                 SpeciesPteropus pumilus Miller, 1910 – Little Golden-mantled Flying Fox 
                                                 SpeciesPteropus rayneri Gray, 1870 – Solomons Flying Fox 
                                                 SpeciesPteropus rennelli Troughton, 1929 – Rennell Flying Fox 
                                                 SpeciesPteropus rodricensis Dobson, 1878 – Rodriguez Flying Fox, Rodrigues Fruit Bat, Rodrigues Island Flying Fox, Rodrigues Flying Fox 
                                                 SpeciesPteropus rufus E. Geoffroy, 1803 – Madagascan Flying Fox, Malagasy Flying Fox 
                                                 SpeciesPteropus samoensis Peale, 1848 – Samoa Flying Fox, Samoan Flying Fox, Common Samoan Flying Fox 
                                                 SpeciesPteropus scapulatus Peters, 1862 – Little Red Flying Fox 
                                                 SpeciesPteropus seychellensis A. Milne-Edwards, 1878 – Seychelles Flying Fox 
                                                 SpeciesPteropus speciosus K. Andersen, 1908 – Philippine Gray Flying Fox 
                                                 SpeciesPteropus subniger (Kerr, 1792) – Dark Flying Fox 
                                                 SpeciesPteropus temminckii Peters, 1867 – Temminck's Flying Fox 
                                                 SpeciesPteropus tokudae Tate, 1934 – Tokuda's Flying Fox, Guam Flying Fox, Little Mariana Fruit Bat 
                                                 SpeciesPteropus tonganus Quoy and Gaimard, 1830 – Insular Flying Fox, Pacific Flying Fox 
                                                 SpeciesPteropus tuberculatus Peters, 1869 – Vanikoro Flying Fox 
                                                 SpeciesPteropus ualanus Peters, 1883 – Kosrae Flying Fox 
                                                 SpeciesPteropus vampyrus (Linnaeus, 1758) – Large Flying Fox 
                                                 SpeciesPteropus vetula Jouan, 1863 – New Caledonia Flying Fox 
                                                 SpeciesPteropus voeltzkowi Matschie, 1909 – Pemba Flying Fox 
                                                 SpeciesPteropus woodfordi Thomas, 1888 – Dwarf Flying Fox 

  Expert: Alfred L. Gardner   
  Notes: Curator of North American mammals and Chief of Mammal Section, National Biological Service, Smithsonian Institution, National Museum of Natural History, Washington, DC, USA   
  Reference for: Pteropus    
  Expert: Don E. Wilson   
  Notes: Smithsonian Institution, MRC 108 Washington DC 20560-0108   
  Reference for: Pteropus    
  Other Source(s):    
  Source: Checklist of CITES species: Index of CITES species, document (version 2013)  
  Acquired: 2013   
  Notes: UNEP-WCMC (Comps.) 2013. Checklist of CITES species. CITES Secretariat, Geneva, Switzerland, and UNEP-WCMC, Cambridge, United Kingdom. Available online at   
  Reference for: Pteropus   
  Source: NODC Taxonomic Code, database (version 8.0)  
  Acquired: 1996   
  Reference for: Pteropus   
  Author(s)/Editor(s): Banks, R. C., R. W. McDiarmid, and A. L. Gardner   
  Publication Date: 1987   
  Article/Chapter Title: Checklist of Vertebrates of the United States, the U.S. Territories, and Canada   
  Journal/Book Name, Vol. No.: Resource Publication, no. 166   
  Page(s): 79   
  Publisher: United States Department of the Interior Fish and Wildlife Service   
  Publication Place: Washington, D.C., USA   
  Reference for: Pteropus   
  Author(s)/Editor(s): Simmons, Nancy B. / Wilson, Don E., and DeeAnn M. Reeder, eds.   
  Publication Date: 2005   
  Article/Chapter Title: Order Chiroptera   
  Journal/Book Name, Vol. No.: Mammal Species of the World: A Taxonomic and Geographic Reference, 3rd ed., vols. 1 & 2   
  Page(s): 312-529   
  Publisher: Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-8221-4   
  Notes: Corrections were made to text at 3rd printing   
  Reference for: Pteropus   
  Author(s)/Editor(s): Wilson, D. E., and R. A. Mittermeier, eds.   
  Publication Date: 2019   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Handbook of the Mammals of the World. Vol. 9: Bats   
  Page(s): 1008   
  Publisher: Lynx Edicions   
  Publication Place: Barcelona, Spain   
  ISBN/ISSN: 978-84-16728190   
  Reference for: Pteropus   
  Author(s)/Editor(s): Wilson, Don E., and DeeAnn M. Reeder, eds.   
  Publication Date: 1993   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Mammal Species of the World: A Taxonomic and Geographic Reference, 2nd ed., 3rd printing   
  Page(s): xviii + 1207   
  Publisher: Smithsonian Institution Press   
  Publication Place: Washington, DC, USA   
  ISBN/ISSN: 1-56098-217-9   
  Notes: Corrections were made to text at 3rd printing   
  Reference for: Pteropus   

 Geographic Information
  Geographic Division:    



  Comment: Comments: The International Commission on Zoological Nomenclature ruled in favor of rejecting Brisson (1762), but conserved several of the generic names including Pteropus (Opinion 1894, ICZN 1998). Formerly included arquatus and leucotis which were transferred to Acerodon by Musser et al. (1982a). Species groups follow Koopman (1994)  
    CITES: Appendix II except Pteropus brunneus and the species included in Appendix I, per 2013 Checklist of CITES species: Index of CITES species published with amendments to the Appendices and changes in standard scientific nomenclature following the 16th meeting of the Conference of the Parties (Bangkok, 2013). For current status see  



Disclaimer: ITIS taxonomy is based on the latest scientific consensus available, and is provided as a general reference source for interested parties. However, it is not a legal authority for statutory or regulatory purposes. While every effort has been made to provide the most reliable and up-to-date information available, ultimate legal requirements with respect to species are contained in provisions of treaties to which the United States is a party, wildlife statutes, regulations, and any applicable notices that have been published in the Federal Register. For further information on U.S. legal requirements with respect to protected taxa, please contact the U.S. Fish and Wildlife Service.

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