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Publication: Savage, Harry M., R. Wills Flowers, and Wendy Porras V. 2005 Rediscovery of Choroterpes atramentum in Costa Rica, type species of Tikuna new genus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae), and its role in the "Great American Interchange" Zootaxa, no. 932 1-14
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