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Publication: Helgen, K. M., F. R. Cole, L. E. Helgen, and D. E. Wilson 2009 Generic revision in the Holarctic ground squirrel genus Spermophilus Journal of Mammalogy, vol. 90, no. 2 270-305

ITIS records citing this publication
 
 Kingdom Animalia
  Anisonyx Rafinesque, 1817 -- invalid
  Callospermophilus Merriam, 1897 -- valid -- Golden-mantled Ground Squirrels
  Callospermophilus lateralis (Say, 1823) -- valid -- Golden-mantled Ground Squirrel
  Callospermophilus lateralis arizonensis (V. Bailey, 1913) -- valid
  Callospermophilus lateralis bernardinus (Merriam, 1898) -- valid
  Callospermophilus lateralis caryi A. H. Howell, 1917 -- invalid
  Callospermophilus lateralis castanurus (Merriam, 1890) -- valid
  Callospermophilus lateralis certus (Goldman, 1921) -- valid
  Callospermophilus lateralis chrysodeirus (Merriam, 1890) -- valid
  Callospermophilus lateralis cinerascens (Merriam, 1890) -- valid
  Callospermophilus lateralis connectens (A. H. Howell, 1931) -- valid
  Callospermophilus lateralis lateralis (Say, 1823) -- valid
  Callospermophilus lateralis mitratus (A. H. Howell, 1931) -- valid
  Callospermophilus lateralis tescorum (Hollister, 1911) -- valid
  Callospermophilus lateralis trepidus (Taylor, 1910) -- valid
  Callospermophilus lateralis trinitatus (Merriam, 1901) -- valid
  Callospermophilus madrensis (Merriam, 1901) -- valid -- Sierra Madre Ground Squirrel
  Callospermophilus saturatus (Rhoads, 1895) -- valid -- Cascade Golden-mantled Ground Squirrel
  Citellus variegatus juglans V. Bailey, 1913 -- invalid
  Citillus Lichtenstein, 1830 -- invalid
  Colobates Milne-Edwards, 1874 -- invalid
  Ictidomys J. A. Allen, 1877 -- valid -- lined ground squirrels
  Ictidomys mexicanus (Erxleben, 1777) -- valid -- Mexican Ground Squirrel, Ardillón mexicano
  Ictidomys parvidens (Mearns, 1896) -- valid -- Rio Grande Ground Squirrel
  Ictidomys tridecemlineatus (Mitchill, 1821) -- valid -- Thirteen-lined Ground Squirrel
  Ictidomys tridecemlineatus alleni (Merriam, 1898) -- valid -- Allen's 13-lined ground squirrel
  Ictidomys tridecemlineatus arenicola (A. H. Howell, 1928) -- valid
  Ictidomys tridecemlineatus blanca (Armstrong, 1971) -- valid
  Ictidomys tridecemlineatus hollisteri (V. Bailey, 1913) -- valid
  Ictidomys tridecemlineatus monticola (A. H. Howell, 1928) -- valid -- White Mountains ground squirrel
  Ictidomys tridecemlineatus olivaceus (J. A. Allen, 1895) -- valid
  Ictidomys tridecemlineatus pallidus (J. A. Allen, 1874) -- valid
  Ictidomys tridecemlineatus parvus (J. A. Allen, 1895) -- valid
  Ictidomys tridecemlineatus texensis (Merriam, 1898) -- valid
  Ictidomys tridecemlineatus tridecemlineatus (Mitchill, 1821) -- valid
  Notocitellus A. H. Howell, 1938 -- valid -- tropical ground squirrels
  Notocitellus adocetus (Merriam, 1903) -- valid -- Tropical Ground Squirrel
  Notocitellus adocetus adocetus (Merriam, 1903) -- valid
  Notocitellus adocetus infernatus (Alvarez and Ramírez-P., 1968) -- valid
  Notocitellus annulatus (Audubon and Bachman, 1842) -- valid -- Ring-tailed Ground Squirrel
  Notocitellus annulatus annulatus (Audubon and Bachman, 1842) -- valid
  Notocitellus annulatus goldmani (Merriam, 1902) -- valid
  Otocolobus Brandt, 1844 -- invalid
  Otospermophilus Brandt, 1844 -- valid -- rock squirrels
  Otospermophilus atricapillus (Bryant, 1889) -- valid -- Baja California Rock Squirrel
  Otospermophilus beecheyi (Richardson, 1829) -- valid -- California Ground Squirrel
  Otospermophilus beecheyi beecheyi (Richardson, 1829) -- valid
  Otospermophilus beecheyi douglasii (Richardson, 1829) -- valid
  Otospermophilus beecheyi fisheri (Merriam, 1893) -- valid
  Otospermophilus beecheyi nesioticus (Elliot, 1904) -- valid
  Otospermophilus beecheyi nudipes (Huey, 1931) -- valid
  Otospermophilus beecheyi parvulus (A. H. Howell, 1931) -- valid
  Otospermophilus beecheyi rupinarum (Huey, 1931) -- valid
  Otospermophilus beecheyi sierrae (A. H. Howell, 1938) -- valid
  Otospermophilus variegatus (Erxleben, 1777) -- valid -- Rock Squirrel
  Otospermophilus variegatus buckleyi (Slack, 1861) -- valid
  Otospermophilus variegatus couchii (Baird, 1855) -- valid
  Otospermophilus variegatus grammurus (Say, 1823) -- valid
  Otospermophilus variegatus robustus (Durrant and Hansen, 1954) -- valid
  Otospermophilus variegatus rupestris (J. A. Allen, 1903) -- valid
  Otospermophilus variegatus tularosae (Benson, 1932) -- valid
  Otospermophilus variegatus utah (Merriam, 1903) -- valid
  Otospermophilus variegatus variegatus (Erxleben, 1777) -- valid
  Poliocitellus A. H. Howell, 1938 -- valid -- Franklin's ground squirrel
  Poliocitellus franklinii (Sabine, 1822) -- valid -- Franklin's Ground Squirrel
  Spermatophilus Wagler, 1830 -- invalid
  Spermophilus F. Cuvier, 1825 -- valid -- Ground Squirrels, old world ground squirrels
  Spermophilus (Spermophilus) F. Cuvier, 1825 -- invalid
  Spermophilus adocetus arceliae (Villa-R., 1942) -- invalid
  Spermophilus altaicus (Eversmann, 1841) -- invalid
  Spermophilus armatus Kennicott, 1863 -- invalid -- Uinta Ground Squirrel
  Spermophilus atricapillus Bryant, 1889 -- invalid -- Baja California Rock Squirrel
  Spermophilus beecheyi (Richardson, 1829) -- invalid -- Ardillón de California, California Ground Squirrel
  Spermophilus beldingi Merriam, 1888 -- invalid -- Belding's Ground Squirrel
  Spermophilus brevicauda Brandt, 1843 -- valid -- Brandt's Ground Squirrel
  Spermophilus brunneus (A. H. Howell, 1928) -- invalid -- northern Idaho ground squirrel, Idaho Ground Squirrel
  Spermophilus buccatus (Lichtenstein, 1830) -- invalid
  Spermophilus canus Merriam, 1898 -- invalid -- Merriam's Ground Squirrel, gray ground squirrel
  Spermophilus citellus (Linnaeus, 1766) -- valid -- European Ground Squirrel, European Souslik
  Spermophilus citellus citellus (Linnaeus, 1766) -- valid
  Spermophilus citellus gradojevici (Martino, 1929) -- valid
  Spermophilus citellus istricus (Calinescu, 1934) -- valid
  Spermophilus citellus martinoi (Peshchev, 1955) -- valid
  Spermophilus columbianus (Ord, 1815) -- invalid -- Columbian Ground Squirrel
  Spermophilus dauricus Brandt, 1843 -- valid -- Daurian Ground Squirrel
  Spermophilus elegans Kennicott, 1863 -- invalid -- Wyoming Ground Squirrel
  Spermophilus erythrogenys Brandt, 1841 -- valid -- Red-cheeked Ground Squirrel
  Spermophilus fulvus (Lichtenstein, 1823) -- valid -- Yellow Ground Squirrel
  Spermophilus fulvus fulvus (Lichtenstein, 1823) -- valid
  Spermophilus fulvus hypoleucos (Satunin, 1909) -- valid
  Spermophilus fulvus oxianus (Thomas, 1915) -- valid
  Spermophilus lateralis (Say, 1823) -- invalid -- Golden-mantled Ground Squirrel
  Spermophilus macedonicus Fraguedakis-Tsolis and Ondrias, 1977 -- invalid
  Spermophilus macrourus Bennett, 1833 -- invalid
  Spermophilus madrensis (Merriam, 1901) -- invalid -- Sierra Madre Ground Squirrel
  Spermophilus major (Pallas, 1778) -- valid -- Russet Ground Squirrel
  Spermophilus mexicanus (Erxleben, 1777) -- invalid -- Ardillón mexicano, Mexican Ground Squirrel
  Spermophilus mexicanus mexicanus (Erxleben, 1777) -- invalid
  Spermophilus mexicanus parvidens Mearns, 1896 -- invalid
  Spermophilus mohavensis Merriam, 1889 -- invalid -- Mohave Ground Squirrel
  Spermophilus mollis Kennicott, 1863 -- invalid -- Piute Ground Squirrel
  Spermophilus pallidicauda (Satunin, 1903) -- valid -- Pallid Ground Squirrel
  Spermophilus parryii (Richardson, 1825) -- invalid -- Arctic Ground Squirrel
  Spermophilus perotensis Merriam, 1893 -- invalid -- Perote Ground Squirrel
  Spermophilus pygmaeus (Pallas, 1778) -- valid -- Little Ground Squirrel
  Spermophilus pygmaeus brauneri (Martino, V. and E., 1917) -- valid
  Spermophilus pygmaeus herbicolus (Martino, V. and E., 1916) -- valid
  Spermophilus pygmaeus mugosaricus (Lichtenstein, 1823) -- valid
  Spermophilus pygmaeus musicus Ménétries, 1832 -- invalid -- Caucasian mountain ground squirrel
  Spermophilus pygmaeus pygmaeus (Pallas, 1778) -- valid
  Spermophilus ralli (Kuznetsov, 1948) -- valid -- Tien Shan Ground Squirrel
  Spermophilus relictus (Kashkarov, 1923) -- valid -- Relict Ground Squirrel
  Spermophilus richardsonii (Sabine, 1822) -- invalid -- Richardson's Ground Squirrel
  Spermophilus saturatus (Rhoads, 1895) -- invalid -- Cascade Golden-mantled Ground Squirrel
  Spermophilus spilosoma Bennett, 1833 -- invalid -- Ardillón punteado, Spotted Ground Squirrel
  Spermophilus suslicus boristhenicus (Pusanov, 1958) -- valid
  Spermophilus suslicus guttatus (Pallas, 1770) -- valid
  Spermophilus suslicus suslicus (Güldenstaedt, 1770) -- valid
  Spermophilus taurensis Gündüz, Jaarola, Tez, Yeniyurt, Polly, and Searle, 2007 -- valid -- Taurus Ground Squirrel
  Spermophilus tereticaudus Baird, 1858 -- invalid -- Ardillón cola redonda, Round-tailed Ground Squirrel
  Spermophilus tereticaudus chlorus (Elliot, 1904) -- invalid -- Coachella Valley round-tailed ground squirrel
  Spermophilus tereticaudus eremonomus (Elliot, 1904) -- invalid
  Spermophilus tereticaudus tereticaudus Baird, 1858 -- invalid
  Spermophilus tereticaudus vociferans (Huey, 1926) -- invalid
  Spermophilus townsendii Bachman, 1839 -- invalid -- Townsend's Ground Squirrel
  Spermophilus tridecemlineatus (Mitchill, 1821) -- invalid -- Thirteen-lined Ground Squirrel
  Spermophilus tridecemlineatus alleni Merriam, 1898 -- invalid -- Allen's 13-lined ground squirrel
  Spermophilus tridecemlineatus monticola (A. H. Howell, 1928) -- invalid
  Spermophilus tridecemlineatus olivaceous (J. A. Allen, 1895) -- invalid
  Spermophilus undulatus (Pallas, 1778) -- invalid -- Long-tailed Ground Squirrel
  Spermophilus variegatus (Erxleben, 1777) -- invalid -- Ardillón de roca, Rock Squirrel
  Spermophilus variegatus tiburonensis Jones and Manning, 1989 -- invalid
  Spermophilus vigilis (Merriam, 1913) -- invalid
  Spermophilus washingtoni (A. H. Howell, 1938) -- invalid -- Washington Ground Squirrel
  Spermophilus washingtoni loringi (A. H. Howell, 1938) -- invalid
  Spermophilus xanthoprymnus (Bennett, 1835) -- valid -- Asia Minor Ground Squirrel
  Urocitellus Obolenskij, 1927 -- valid -- Holarctic ground squirrels
  Urocitellus armatus (Kennicott, 1863) -- valid -- Uinta Ground Squirrel
  Urocitellus beldingi (Merriam, 1888) -- valid -- Belding's Ground Squirrel
  Urocitellus beldingi beldingi (Merriam, 1888) -- valid
  Urocitellus beldingi creber (Hall, 1940) -- valid
  Urocitellus beldingi oregonus (Merriam, 1898) -- valid
  Urocitellus brunneus (A. H. Howell, 1928) -- valid -- Idaho Ground Squirrel
  Urocitellus brunneus brunneus (A. H. Howell, 1928) -- valid -- northern Idaho ground squirrel
  Urocitellus brunneus endemicus (Yensen, 1991) -- valid
  Urocitellus canus (Merriam, 1898) -- valid -- Merriam's Ground Squirrel
  Urocitellus columbianus (Ord, 1815) -- valid -- Columbian Ground Squirrel
  Urocitellus columbianus columbianus (Ord, 1815) -- valid
  Urocitellus columbianus ruficaudus (A. H. Howell, 1928) -- valid
  Urocitellus elegans (Kennicott, 1863) -- valid -- Wyoming Ground Squirrel
  Urocitellus elegans aureus (Davis, 1939) -- valid
  Urocitellus elegans elegans (Kennicott, 1863) -- valid
  Urocitellus elegans nevadensis (A. H. Howell, 1928) -- valid
  Urocitellus mollis (Kennicott, 1863) -- valid -- Piute Ground Squirrel
  Urocitellus mollis artemesiae (Merriam, 1913) -- valid
  Urocitellus mollis idahoensis (Merriam, 1913) -- valid
  Urocitellus mollis mollis (Kennicott, 1863) -- valid
  Urocitellus parryii (Richardson, 1825) -- valid -- Arctic Ground Squirrel
  Urocitellus parryii ablusus (Osgood, 1903) -- valid
  Urocitellus parryii kennicottii (Ross, 1861) -- valid
  Urocitellus parryii kodiacensis (J. A. Allen, 1874) -- valid
  Urocitellus parryii leucostictus (Brandt, 1844) -- valid
  Urocitellus parryii lyratus (Hall and Gilmore, 1932) -- valid
  Urocitellus parryii nebulicola (Osgood, 1903) -- valid
  Urocitellus parryii osgoodi (Merriam, 1900) -- valid
  Urocitellus parryii parryii (Richardson, 1825) -- valid
  Urocitellus parryii plesius (Osgood, 1900) -- valid
  Urocitellus parryii stejnegeri (J. A. Allen, 1903) -- valid
  Urocitellus richardsonii (Sabine, 1822) -- valid -- Richardson's Ground Squirrel
  Urocitellus townsendii (Bachman, 1839) -- valid -- Townsend's Ground Squirrel
  Urocitellus townsendii nancyae (Nadler, 1968) -- valid
  Urocitellus townsendii townsendii (Bachman, 1839) -- valid
  Urocitellus undulatus (Pallas, 1778) -- valid -- Long-tailed Ground Squirrel
  Urocitellus undulatus jacutensis (Brandt, 1844) -- valid
  Urocitellus undulatus menzbieri (Ognev, 1937) -- valid
  Urocitellus undulatus stramineus (Obolenskii, 1927) -- valid
  Urocitellus undulatus transbaikalicus (Obolenskii, 1927) -- valid
  Urocitellus undulatus undulatus (Pallas, 1778) -- valid
  Urocitellus washingtoni (A. H. Howell, 1938) -- valid -- Washington Ground Squirrel
  Xerospermophilus Merriam, 1892 -- valid -- pygmy ground squirrels
  Xerospermophilus mohavensis (Merriam, 1889) -- valid -- Mohave Ground Squirrel
  Xerospermophilus perotensis (Merriam, 1893) -- valid -- Perote Ground Squirrel
  Xerospermophilus spilosoma (Bennett, 1833) -- valid -- Spotted Ground Squirrel
  Xerospermophilus spilosoma altiplanensis (Anderson, 1972) -- valid
  Xerospermophilus spilosoma ammophilus (Hoffmeister, 1959) -- valid
  Xerospermophilus spilosoma annectens (Merriam, 1893) -- valid
  Xerospermophilus spilosoma bavicorensis (Anderson, 1972) -- valid
  Xerospermophilus spilosoma cabrerai (Dalquest, 1951) -- valid
  Xerospermophilus spilosoma canescens (Merriam, 1890) -- valid
  Xerospermophilus spilosoma cryptospilotus (Merriam, 1890) -- valid
  Xerospermophilus spilosoma marginatus (V. Bailey, 1902) -- valid
  Xerospermophilus spilosoma obsoletus (Kennicott, 1863) -- valid
  Xerospermophilus spilosoma oricolus (Alvarez, 1962) -- valid
  Xerospermophilus spilosoma pallescens (A. H. Howell, 1928) -- valid
  Xerospermophilus spilosoma pratensis (Merriam, 1890) -- valid
  Xerospermophilus spilosoma spilosoma (Bennett, 1833) -- valid
  Xerospermophilus tereticaudus (Baird, 1858) -- valid -- Round-tailed Ground Squirrel
  Xerospermophilus tereticaudus apricus (Huey, 1927) -- valid
  Xerospermophilus tereticaudus chlorus (Elliot, 1904) -- valid -- Coachella Valley round-tailed ground squirrel, Palm Springs round-tailed ground squirrel
  Xerospermophilus tereticaudus neglectus (Merriam, 1889) -- valid
  Xerospermophilus tereticaudus tereticaudus (Baird, 1858) -- valid
   

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