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Publication: Ota, Hidetoshi, Akiko Ito, and Jun-Tsong Lin 1999 Systematic Review of the Elapid Snakes Allied to Hemibungarus japonicus (Günther, 1868) in the East Asian Islands, with Description of a New Subspecies from the Central Ryukyus Journal of Herpetology, vol. 33, no. 4 675-687
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